Bluff Downs giant python: Difference between revisions

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The '''Bluff Downs giant python''' ('''''Liasis dubudingala''''') is an [[extinction|extinct]] [[species]] of [[snake]] from [[Queensland]], [[Australia]], that lived during the Early [[Pliocene]]. Named in 2002, ''Liasis dubudingala'' was likely the biggest snake found in Australia, with a total length of up to {{convert|9|m|abbr=on}}. This length rivals the largest extant snake species, the [[reticulated python]] from Asia and the [[green anaconda]] from South America. It may have fed on larger prey such as juvenile [[Diprotodontidae|diprotodontids]], but it is also possible that it was a skilled climber capable of catching birds and arboreal marsupials.
 
==History and naming==
The first fossil remains of ''Liasis dubudingala'' were discovered in 1975 by Australian paleontologist [[Mike Archer (paleontologist)|Mike Archer]] in the early Pliocene sediments of the [[Allingham Formation]], [[Queensland]]. The trunk vertebra was initially assigned to the genus ''[[Morelia]]'' by Archer, but later research found sufficient evidence to assign the material to the genus ''Liasis'', with Scanlon and Mackness errectingerecting a new species for the material in 2002. The trunk vertebra discovered by Archer, QMF 9132, was chosen to be the [[holotype]], but a number of additional fossils including vertebrae, ribs, teeth and a dentary were also referred to the species.<ref name=SM01>{{Cite journal | doi = 10.1080/03115510108619232| title = A new giant python from the Pliocene Bluff Downs Local Fauna of northeastern Queensland| journal = Alcheringa: An Australasian Journal of Palaeontology| volume = 25| issue = 4| pages = 425| year = 2001| last1 = Scanlon | first1 = J. D. | last2 = MacKness | first2 = B. S. | s2cid = 85185368}}</ref>
 
The species name "dubudingala" is a combination of the words dubu ("ghost") and dingal ("to squeeze"), both deriving from the [[Guugu Yalandji language]]. This combination, effectively meaning "ghost squeezer", was chosen to reflect the likely constricting habits of the python.<ref name=SM01/> The [[Australian Museum]] furthermore coined the vernacular name Bluff Downs giant python.<ref name=AMW>{{cite web |url=https://web.archive.org/web/20090217125654/http://lostkingdoms.com/facts/factsheet44.htm |title=Australia's Lost Kingdom: Bluff Downs Giant Python |author=<!--Not stated--> |year=2004 |website= Australian Museum|archive-url=https://web.archive.org/web/20090217125654/http://lostkingdoms.com/facts/factsheet44.htm |access-date=August 8, 2022|archive-date=2009-02-17 }}</ref>
 
==Description==
The neural spines of ''Liasis dubudingala'' were higher than they were long and slope backwards, however the dorsoposterior angle of said slope is blunt rather than accute. In general the entire vertebra is notably higher than long, with the neural spine, neural arch and centrum all contributing to the height in equal measures. The roof of the [[zygosphene]] slopes at a steep angle and its junction with the neural spine is blunt. The [[articular processes]] are almost horizontal with a slope of less than 5%. The hypapophysis below the centrum grows gradually more shallow from the front to the back, rather than doing so more abruptly. Additionally, the haemal keel of ''Liasis dubudingala'' is pronounced throughout the trunk of the vertebrae. While the holotype vertebra is from the middle of the snake's trunk, the most numerous vertebrae correspond with the anterior trunk region, before the heart. These precardiac vertebrae are higher than they are long like those of the middle trunk, however to a much greater degree, expressing a ratio of almost 2:1 between height and length. This ratio is less well developed in the posterior trunk vertebrae, which also display longer vertebral centra and less horizontal zygapophyses. At least one vertebra, alongside a dentary fragment, appear to belong to a juvenile specimen. The dentary preserves a large portion of the posterior tooth row, showing ten [[alveoli]] situated on a raised ridge. Safe for the second, fourth and sixth of these alveoli, all teeth in the dentary fragment are [[Tooth ankylosis|ankylosed]], meaning they are permanently fused to the jaw bone. The curvature is similar to those of the modern [[olive python]], although the teeth are smaller in diameter.<ref name=SM01/>
 
''Liasis dubudingala'' was likely among the largest snakes to have existed in Australia, with the largest vertebra meassuringmeasuring a total of {{convert|59.1|mm|abbr=on}} wide. Using Scanlon's 1993 method of estimating the size of fossil snakes, this would result in a total length of {{convert|8.35|m|abbr=on}}. However, this method is based on the proportions of the extant [[Black-headed python]], which is known to have a spine composed of 394 individual vertebrae, contrasting with the average of 450 vertebrae that make up the skeleton of large ''Liasis'' species. This would increase the length of ''Liasis dubudingala'' to roughly {{convert|9|m|abbr=on}}. With this length the Bluff Downs giant python would have been the largest snake of Australia, even exceeding the [[Madtsoiidae|madtsoiid]] snake ''[[Yurlunggur]]''.<ref name=SM01/><ref name=S04>{{Cite journal|last1=Scanlon|first1=J.D.|year=2004|title=First known axis vertebra of a madtsoiid snake (Yurlunggur camfieldensis) and remarks on the neck of snakes.|url=https://www.researchgate.net/profile/John-Scanlon/publication/252044827_First_known_axis_vertebra_of_a_madtsoiid_snake_Yurlunggur_camfieldensis_and_remarks_on_the_neck_of_snakes/links/54015c410cf23d9765a494de/First-known-axis-vertebra-of-a-madtsoiid-snake-Yurlunggur-camfieldensis-and-remarks-on-the-neck-of-snakes.pdf252044827|journal=Beagle: Records of the Museums and Art Galleries of the Northern Territory|volume=20|pages=207207–215|doi=10.5962/p.286329 |s2cid=128423678 |doi-215access=free}}</ref>
 
==Classification==
In the type description of ''Liasis dubudingala'', Scanlon and MacKnees consider the genera ''[[Antaresia]]'', ''[[Bothrochilus]]'', ''[[Leiopython]]'' and ''[[Apodora]]'' to all be synonymous with ''[[Liasis]]''. The authors subsequently diagnose the genus based on a variety of traits including the anatomy of the skull, overall proportions, scales and the anatomy of the [[hemipenis]]. Although none of the traits used to define ''Liasis [[sensu lato]]'' in this manner could be observed in the Bluff Downs giant python, the authors nevertheless assign it to the genus based on general similarities. While the high neural spine can also be found in ''[[Morelia]]'' and ''[[Python (genus)|Python]]'', the Bluff Downs giant python was found to lack several derived traits that would connect it to the afforementionedaforementioned genera. Outside of ''Morelia'' and ''Python'', high neural spines are also found in the species of ''Liasis [[sensu stricto]]'', meaning ''Liasis'' species in the strictest definition (''[[Liasis olivacea]]'' and ''[[Liasis mackloti]]''). Scanlon and MacKnees conclude that based on the available characteristics, ''Liasis dubudingala'' was likely either a sister taxon to or part of ''Liasis sensu stricto''.<ref name=SM01/>
 
==Paleoecology==
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{{Reflist}}
 
{{Fossil snakes}}
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