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{{Year nav topic20|2019|reptile paleontology|science |paleobotany |paleontology |arthropod paleontology |paleoentomology |paleomalacology |archosaur paleontology |mammal paleontology |paleoichthyology}}
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This '''list of [[fossil]] [[reptile]]s described in 2019''' is a list of new [[taxa]] of fossil reptiles that were [[binomial nomenclature|described]] during the year 2019, as well as other significant discoveries and events related to reptile [[paleontology]] that occurred in 2019.
{{Year in paleontology header}}
 
==Lizards and snakes==
===Research===
* A study aiming to test which ecological and developmental traits have influenced skull evolution in the [[Squamata|squamate]] reptiles, based on data from extant and extinct taxa, is published by Watanabe ''et al.'' (2019).<ref>{{Cite journal|author1=Akinobu Watanabe |author2=Anne-Claire Fabre |author3=Ryan N. Felice |author4=Jessica A. Maisano |author5=Johannes Müller |author6=Anthony Herrel |author7=Anjali Goswami |year=2019 |title=Ecomorphological diversification in squamates from conserved pattern of cranial integration |journal=Proceedings of the National Academy of Sciences of the United States of America |volume=116 |issue=29 |pages=14688–14697 |doi=10.1073/pnas.1820967116 |pmid=31262818 |pmc=6642379 |doi-access=free }}</ref>
* A new assemblage of lizard tracks, representing the largest such assemblage yet reported from the [[Cretaceous]], is described from the [[Lower Cretaceous]] [[Jinju Formation]] ([[South Korea]]) by Kim ''et al.'' (2019), who name a new [[ichnotaxon]] ''[[Neosauroides]] innovatus''.<ref>{{Cite journal|author1=Kyung Soo Kim |author2=Jong Deock Lim |author3=Martin G. Lockley |author4=Dong Hee Kim |author5=Laura Piñuela |author6=Jae Sang Yoo |year=2019 |title=Largest Cretaceous lizard track assemblage, new morphotypes and longest trackways comprise diverse components of an exceptional Korean Konservat-Lagerstätten ichnofauna |journal=Scientific Reports |volume=9 |issue=1 |article-number=13278 |doi=10.1038/s41598-019-49442-0 |pmid=31527673 |pmc=6746761 |bibcode=2019NatSR...913278K }}</ref>
* Description of fossils of [[amphisbaenia]]ns and [[Anguimorpha|anguimorph]] lizards from the late [[Miocene]] Solnechnodolsk locality (southern European [[Russia]]) will be published by Černanský, Syromyatnikova & Jablonski (2019).<ref>{{cite journal |author1=Andrej Černanský |author2=Elena V. Syromyatnikova |author3=Daniel Jablonski |year=2019 |title=The first record of amphisbaenian and anguimorph lizards (Reptilia, Squamata) from the upper Miocene Solnechnodolsk locality in Russia |journal=Historical Biology: An International Journal of Paleobiology |volume=in press |issue= |pages= 1–11|doi=10.1080/08912963.2018.1539973 }}</ref>
* A juvenileNew specimen of ''[[EolacertaYabeinosaurus]] robustarobustus'', ispreserving describedtraces fromof theintegument [[Eocene]]and [[Messelabdominal pit]]contents, ([[Germany]])is described by ČerňanskýXing &''et Smithal.'' (2019).<ref>{{citeCite journal |author1=AndrejLida ČerňanskýXing |author2=KristerKechung Niu |author3=Rod TS. SmithTaylor |author4=Susan E. Evans |year=2019 |title=TheIntegumentary firstremains juvenileand specimenabdominal ofcontents in the Early Cretaceous Chinese lizard, ''EolacertaYabeinosaurus'' (Squamata: Eolacertidae), fromdemonstrate thecolour early–middlebanding Eoceneand ofa thediet Messelincluding Pit (Germany)crayfish |journal=ComptesCretaceous Rendus PalevolResearch |volume=in press108 |pagesarticle-number=104320 |doi=10.1016/j.crpvcretres.2019.04104320 |s2cid=210262098 |url=https://discovery.ucl.ac.uk/id/eprint/10089773/1/Yabeinosaurus%20MS.004pdf }}</ref>
* DescriptionJaw elements of late [[Mioceneskink]]s with an anatomy identical to ''[[Lacertidae|lacertidHeremites]] fossilsvittatus'' are described from the Solnechnodolsklate [[Miocene]] locality Solnechnodolsk ([[Russia]]), belongingby toČerňanský a& relativeSyromyatnikova of(2019), extantrepresenting first [[Balkan green lizardNeogene]], isrecord publishedof bythe Čerňanskýclade &[[Mabuyidae]] Syromyatnikovareported (2019)so far.<ref>{{Cite journal|author1=Andrej Čerňanský |author2=Elena V. Syromyatnikova |year=2019 |title=The first Miocenepre-Quaternary fossil fossilsrecord of ''Lacerta''the cf.clade ''trilineata'' (Squamata, Lacertidae)Mabuyidae with a comparativecomment studyon the enclosure of the mainMeckelian cranial osteological differencescanal in green lizards and their relativesskinks |journal=PLoSPapers ONEin Palaeontology |volume=147 |issue=81 |pages=e0216191195–215 |doi=10.13711002/journal.ponespp2.02161911279 |pmidissn=314338072056-2799 |pmcs2cid=6703700204266535 }}</ref>
* A studyjuvenile onspecimen theof diet,''[[Eolacerta]] habitatrobusta'' andis timingdescribed andfrom causethe of[[Eocene]] extinction[[Messel ofpit]] ''([[Gallotia goliathGermany]]'' is published) by CrowleyČerňanský ''et& al.''Smith (2019).<ref>{{cite journal |author1=BrookeAndrej Erin CrowleyČerňanský |author2=YurenaKrister YanesT. |author3=Stella Grace Mosher |author4=Juan Carlos RandoSmith |year=2019 |title=RevisitingThe thefirst foragingjuvenile ecologyspecimen andof extinction''Eolacerta'' history(Squamata: ofEolacertidae) twofrom endemicthe vertebratesearly–middle fromEocene Tenerife,of Canarythe IslandsMessel Pit (Germany) |journal=QuaternaryComptes Rendus Palevol |volume=218 |issue=17 |pages=Article 10735–745 |doi=10.33901016/quat2010010j.crpv.2019.04.004 |s2cid=199092977 }}</ref>
* The first fossilDescription of alate [[fringe-toedMiocene]] lizard[[Lacertidae|lacertid]] reportedfossils sofrom farthe isSolnechnodolsk described from thelocality ([[MioceneRussia]]), [[Dovebelonging Springto Formation]]a relative of extant ([[CaliforniaBalkan green lizard]], [[Unitedis States]])published by ScarpettaČerňanský & Syromyatnikova (2019).<ref>{{citeCite journal|author1=Andrej Čerňanský |authorauthor2=SimonElena GV. ScarpettaSyromyatnikova |year=2019 |title=The first knownMiocene fossilfossils of ''UmaLacerta'': ecologicalcf. evolution''trilineata'' and(Squamata, theLacertidae) originswith a comparative study of Norththe Americanmain fringe-toedcranial osteological differences in green lizards and their relatives |journal=BMC EvolutionaryPLOS BiologyONE |volume=1914 |issue=8 |pages=Article number 178e0216191 |doi=10.11861371/s12862-019journal.pone.0216191 |pmid=31433807 |pmc=6703700 |bibcode=2019PLoSO..1416191C |doi-1501-5access=free }}</ref>
* A study on the taxonomydiet, habitat and evolutionarytiming historyand cause of [[Glyptosaurinae|glyptosaurine]]extinction of ''[[Anguidae|anguidsGallotia goliath]]'' is published by SullivanCrowley ''et al.'' (2019).<ref>{{cite journal |authorauthor1=RobertBrooke M.Erin Crowley |author2=Yurena Yanes |author3=Stella Grace Mosher |author4=Juan Carlos SullivanRando |year=2019 |title=TheRevisiting taxonomy,the chronostratigraphyforaging ecology and paleobiogeographyextinction history of glyptosaurinetwo lizardsendemic (Glyptosaurinaevertebrates from Tenerife, Anguidae)Canary Islands |journal=Comptes Rendus PalevolQuaternary |volume=in press2 |issue=1 |pagesarticle-number=10 |doi=10.10163390/j.crpv.2019.05.006quat2010010 |doi-access=free }}</ref>
* An almost complete and articulated fossil skull of the [[La Palma giant lizard]] is described from the [[Pleistocene]] of the [[La Palma]] island ([[Canary Islands]]) by Cruzado-Caballero ''et al.'' (2019), who also study the phylogenetic relationships of extant and fossil members of the genus ''[[Gallotia]]''.<ref>{{Cite journal|author1=Penélope Cruzado-Caballero |author2=Carolina Castillo Ruiz |author3=Arnau Bolet |author4=Juan Ramón Colmenero |author5=Julio De la Nuez |author6=Ramón Casillas |author7=Sergio Llacer |author8=Federico Bernardini |author9=Josep Fortuny |year=2019 |title=First nearly complete skull of ''Gallotia auaritae'' (lower-middle Pleistocene, Squamata, Gallotiinae) and a morphological phylogenetic analysis of the genus ''Gallotia'' |journal=Scientific Reports |volume=9 |issue=1 |article-number=16629 |doi=10.1038/s41598-019-52244-z |pmid=31719546 |pmc=6851374 |bibcode=2019NatSR...916629C }}</ref>
* Fossils of ''[[Peltosaurus]] granulosus'' are described from the middle [[Oligocene]] [[Monroe Creek Formation|Monroe Creek]] and upper [[Sharps Formation|Sharps]] formations of Sharps Corner ([[South Dakota]], [[United States]]) by Scarpetta (2019), representing the youngest known record of glyptosaurine lizards reported so far.<ref>{{Cite journal|author=Simon G. Scarpetta |year=2019 |title=''Peltosaurus granulosus'' (Squamata, Anguidae) from the middle Oligocene of Sharps Corner, South Dakota, and the youngest known chronostratigraphic occurrence of Glyptosaurinae |journal=Journal of Vertebrate Paleontology |volume=in press |issue= |pages=e1622129 |doi=10.1080/02724634.2019.1622129 }}</ref>
* FossilThe first fossil of a [[Anguidae|anguinefringe-toed lizard]] materialreported so far is described from the lower [[Miocene]] locality[[Dove UlmSpring – WesttangenteFormation]] ([[GermanyCalifornia]]), for[[United the first timeStates]]) by Klembara, Hain & ČerňanskýScarpetta (2019).<ref>{{cite journal |author1author=JozefSimon Klembara |author2=Miroslav Hain |author3=AndrejG. ČerňanskýScarpetta |year=2019 |title=The first recordknown offossil anguine''Uma'': lizardsecological (Anguimorpha,evolution Anguidae) fromand the earlyorigins Mioceneof localityNorth UlmAmerican fringe-toed Westtangente in Germanylizards |journal=HistoricalBMC Evolutionary Biology: An International Journal of Paleobiology |volume=3119 |issue=81 |pagesarticle-number=1016–1027178 |doi=10.10801186/08912963.2017.1416469s12862-019-1501-5 |pmid=31492110 |pmc=6729053 |doi-access=free }}</ref>
* A study on the taxonomy and evolutionary history of [[Glyptosaurinae|glyptosaurine]] [[Anguidae|anguids]] is published by Sullivan (2019).<ref>{{cite journal |author=Robert M. Sullivan |year=2019 |title=The taxonomy, chronostratigraphy and paleobiogeography of glyptosaurine lizards (Glyptosaurinae, Anguidae) |journal=Comptes Rendus Palevol |volume=18 |issue=7 |pages=747–763 |doi=10.1016/j.crpv.2019.05.006 |s2cid=202174449 }}</ref>
* Bochaton ''et al.'' (2019) present equations producing estimations of size and weight of [[monitor lizard]]s on the basis of measurements of their bones, and use them to determine body size and weight of Late Pleistocene monitor lizards known from remains from the Doi Pha Kan rock shelter ([[Thailand]]).<ref>{{cite journal |author1=Corentin Bochaton |author2=Pauline Hanot |author3=Stéphane Frère |author4=Julien Claude |author5=Wilailuck Naksri |author6=Prasit Auetrakulvit |author7=Valéry Zeitoun |year=2019 |title=Size and weight estimations of subfossil monitor lizards (''Varanus'' sp. Merrem 1820) with an application to the Hoabinhian assemblage of Doi Pha Kan (Late Pleistocene, Lampang province, Thailand) |journal=Annales de Paléontologie |volume=in press |pages= |doi=10.1016/j.annpal.2019.05.003 }}</ref>
* Fossils of ''[[Peltosaurus]] granulosus'' are described from the middle [[Oligocene]] [[Monroe Creek Formation|Monroe Creek]] and upper [[Sharps Formation|Sharps]] formations of [[Sharps Corner, South Dakota]] by Scarpetta (2019), representing the youngest known record of glyptosaurine lizards reported so far.<ref>{{Cite journal|author=Simon G. Scarpetta |year=2019 |title=''Peltosaurus granulosus'' (Squamata, Anguidae) from the middle Oligocene of Sharps Corner, South Dakota, and the youngest known chronostratigraphic occurrence of Glyptosaurinae |journal=Journal of Vertebrate Paleontology |volume=39 |issue=3 |pages=e1622129 |doi=10.1080/02724634.2019.1622129 |s2cid=196690362 }}</ref>
* A study evaluating the fossil record of [[mosasaur]]s in terms of fossil completeness as a measure of fossil quality is published by Driscoll ''et al.'' (2019).<ref>{{Cite journal|author1=Daniel A. Driscoll |author2=Alexander M. Dunhill |author3=Thomas L. Stubbs |author4=Michael J. Benton |year=2019 |title=The mosasaur fossil record through the lens of fossil completeness |journal=Palaeontology |volume=62 |issue=1 |pages=51–75 |doi=10.1111/pala.12381 }}</ref>
* AFossil study[[Anguidae|anguine]] onmaterial theis morphologydescribed offrom the marginal teeth oflower ''[[MosasaurusMiocene]] lemonnieri'',locality andUlm on theirWesttangente implications([[Germany]]) for the distinguishabilityfirst oftime thisby species on the basis of fossil teethKlembara, willHain be published by& MadziaČerňanský (2019).<ref>{{cite journal |authorauthor1=DanielJozef MadziaKlembara |author2=Miroslav Hain |author3=Andrej Čerňanský |year=2019 |title=DentalThe variabilityfirst andrecord distinguishabilityof inanguine ''Mosasaurus lemonnieri''lizards (MosasauridaeAnguimorpha, Anguidae) from the Campanianearly andMiocene Maastrichtianlocality ofUlm Belgium, andWesttangente implications for taxonomic assessments of mosasauridin dentitionsGermany |journal=Historical Biology: An International Journal of Paleobiology |volume=in press31 |issue=8 |pages=1016–1027 1–15|doi=10.1080/08912963.20192017.15888921416469 |s2cid=90251378 }}</ref>
* ABochaton skull''et ofal.'' a(2019) memberpresent ofequations theproducing genusestimations ''Mosasaurus''of issize reportedand fromweight the upperof [[Maastrichtianmonitor lizard]]s [[Lopezon dethe Bertodanobasis Formation]]of ([[Seymourmeasurements Island]]of their bones, [[Antarctica]])and byuse Ruizthem ''etto al.''determine body size and weight of Late Pleistocene monitor lizards known from remains from the Doi Pha Kan rock shelter (2019[[Thailand]]).<ref>{{cite journal |author1=Pablo GonzálezCorentin RuizBochaton |author2=MartaPauline S. FernándezHanot |author3=MarianellaStéphane TaleviFrère |author4=Juan M.Julien LeardiClaude |author5=MarceloWilailuck A.Naksri Reguero|author6=Prasit Auetrakulvit |author7=Valéry Zeitoun |year=2019 |title=ASize newand Plotosauriniweight mosasaurestimations skullof fromsubfossil monitor lizards (''Varanus'' sp. Merrem 1820) with an application to the upperHoabinhian Maastrichtianassemblage of Antarctica.Doi PlotosauriniPha paleogeographicKan occurrences(Late Pleistocene, Lampang province, Thailand) |journal=CretaceousAnnales de ResearchPaléontologie |volume=103105 |issue=4 |pages=Article 104166295–304 |doi=10.1016/j.cretresannpal.2019.0605.012003 |s2cid=201331999 }}</ref>
* A study onevaluating the anatomyfossil of the inner earrecord of ''[[Platecarpusmosasaur]]''s willin beterms of fossil completeness as a measure of fossil quality is published by YiDriscoll &''et [[Mark Norell|Norell]]al.'' (2019).<ref>{{Cite journal|author1=HongyuDaniel YiA. Driscoll |author2=MarkAlexander M. Dunhill |author3=Thomas L. Stubbs |author4=Michael J. NorellBenton |year=2019 |title=The bonymosasaur labyrinthfossil ofrecord ''Platecarpus''through (Squamata:the Mosasauria)lens andof aquatic adaptations infossil squamate reptilescompleteness |journal=PalaeoworldPalaeontology |volume=in press62 |issue=1 |pages=51–75 |doi=10.10161111/jpala.palwor12381 |s2cid=133743666 |url=http://eprints.2018whiterose.12ac.uk/130080/1/Driscoll%20et%20al%20accepted.001pdf }}</ref>
* A study on the morphology of the marginal teeth of ''[[Mosasaurus]] lemonnieri'', and on their implications for the distinguishability of this species on the basis of fossil teeth, is published online by Madzia (2019).<ref>{{cite journal |author=Daniel Madzia |year=2019 |title=Dental variability and distinguishability in ''Mosasaurus lemonnieri'' (Mosasauridae) from the Campanian and Maastrichtian of Belgium, and implications for taxonomic assessments of mosasaurid dentitions |journal=Historical Biology: An International Journal of Paleobiology |volume=32 |issue=10 |pages=1340–1354 |doi=10.1080/08912963.2019.1588892 |s2cid=108526638 }}</ref>
* An isolated tooth of a [[Tylosaurinae|tylosaurine]] mosasaur is described from the [[Turonian]] of the Apennine Carbonate Platform by Romano ''et al.'' (2019), representing the first tylosaurine from [[Italy]] and the southernmost occurrence of a tylosaurine in the northern margin of the Mediterranean [[Tethys Ocean|Tethys]].<ref>{{Cite journal|author1=Marco Romano |author2=Riccardo Manni |author3=Enrico Venditti |author4=Umberto Nicosia |author5=Angelo Cipriani |year=2019 |title=First occurrence of a Tylosaurinae mosasaur from the Turonian of the Central Apennines, Italy |journal=Cretaceous Research |volume=96 |pages=196–209 |doi=10.1016/j.cretres.2019.01.001 }}</ref>
* A studyskull onof thea phylogenetic relationshipsmember of tylosaurinethe genus mosasaurs''Mosasaurus'' is publishedreported from the upper [[Maastrichtian]] [[Lopez de Bertodano Formation]] ([[Seymour Island]], [[Antarctica]]) by Jiménez-HuidobroRuiz &''et Caldwellal.'' (2019).<ref>{{Citecite journal |author1=PaulinaPablo Jiménez-HuidobroGonzález Ruiz |author2=MichaelMarta WS. CaldwellFernández |author3=Marianella Talevi |author4=Juan M. Leardi |author5=Marcelo A. Reguero |year=2019 |title=A new hypothesisPlotosaurini ofmosasaur skull from the phylogeneticupper relationshipsMaastrichtian of theAntarctica. TylosaurinaePlotosaurini (Squamata:paleogeographic Mosasauroidea)occurrences |journal=FrontiersCretaceous in Earth ScienceResearch |volume=7103 |pagesarticle-number=Article 47104166 |doi=10.33891016/feartj.cretres.2019.0004706.012 |s2cid=198418273 |url=https://rid.unrn.edu.ar/jspui/handle/20.500.12049/5094 |hdl=11336/125124 |hdl-access=free }}</ref>
* A reviewstudy on the anatomy of the taxonomicinner historyear of ''[[ClidastesPlatecarpus]] liodontus'' and "Clidastes moorevillensis" is published by LivelyYi & [[Mark Norell|Norell]] (2019).<ref>{{Cite journal|authorauthor1=JoshuaHongyu R.Yi Lively|author2=Mark Norell |year=2019 |title=TaxonomyThe andbony historicallabyrinth inertia:of ''ClidastesPlatecarpus'' (Squamata: MosasauridaeMosasauria) asand aaquatic caseadaptations studyin ofsquamate problematic paleobiological taxonomyreptiles |journal=Alcheringa: An Australasian Journal of PalaeontologyPalaeoworld |volume=in press28 |issue= 4 |pages=550–561 516–527|doi=10.10801016/03115518j.palwor.2018.154968512.001 |s2cid=134163806 }}</ref>
* AAn juvenileisolated mosasaurtooth specimenof affected bya [[infectious arthritisTylosaurinae|tylosaurine]] and [[spondyloarthropathy]]mosasaur is described from the upper [[MaastrichtianTuronian]] of [[Antarctica]]the Apennine Carbonate Platform by TaleviRomano ''et al.'' (2019), representing the first reporttylosaurine offrom a[[Italy]] skeletaland pathologythe southernmost occurrence of a mosasaurtylosaurine fromin the northern margin of the Mediterranean [[SouthernTethys HemisphereOcean|Tethys]].<ref>{{Cite journal|author1=MarianellaMarco TaleviRomano |author2=BruceRiccardo RothschildManni |author3=MartaEnrico FernándezVenditti |author4=MarceloUmberto RegueroNicosia |author5=MatíasAngelo MitidieriCipriani |year=2019 |title=AFirst pathologicaloccurrence scapula inof a Tylosaurinae mosasaur from the upper MaastrichtianTuronian of Antarctica:the evidenceCentral of infectious arthritis andApennines, spondyloarthropathyItaly |journal=Cretaceous Research |volume=10096 |pages=1–4196–209 |doi=10.1016/j.cretres.2019.0301.024001 |s2cid=135342154 }}</ref>
* A study on the evolutionphylogenetic of vertebral intercentrum systemrelationships of snakes,tylosaurine as indicated by data from specimens of ''[[Najash]] rionegrina'' and ''[[Dinilysia]] patagonica'',mosasaurs is published by GarberoglioJiménez-Huidobro ''et& al.''Caldwell (2019).<ref>{{Cite journal|author1=FernandoPaulina F. GarberoglioJiménez-Huidobro |author2=Raúl O. Gómez |author3=Tiago R. Simões |author4=Michael W. Caldwell |author5=Sebastián Apesteguía |year=2019 |title=TheA evolutionnew hypothesis of the axialphylogenetic skeletonrelationships intercentrum system in snakes revealed by new data fromof the CretaceousTylosaurinae snakes(Squamata: ''Dinilysia'' and ''Najash''Mosasauroidea) |journal=ScientificFrontiers in Earth ReportsScience |volume=97 |issuearticle-number=1 |pages=Article number 127647 |doi=10.10383389/s41598-018-36979-9feart.2019.00047 |pmidbibcode=307185252019FrEaS...7...47J |pmcdoi-access=6362196free }}</ref>
* NewA specimenreview of ''Najashthe rionegrina'',taxonomic consistinghistory of a''[[Clidastes]] partial skullliodontus'' and closely"Clidastes associated vertebrae,moorevillensis" is describedpublished by Garberoglio ''et al.''Lively (2019).<ref>{{Cite journal|author1author=FernandoJoshua FR. Garberoglio |author2=Raúl O. Gómez |author3=Sebastián Apesteguía |author4=Michael W. Caldwell |author5=María L. Sánchez |author6=Gonzalo VeigaLively |year=2019 |title=A new specimen with skullTaxonomy and vertebraehistorical ofinertia: ''Najash rionegrinaClidastes'' (LepidosauriaSquamata: OphidiaMosasauridae) fromas thea earlycase Late Cretaceousstudy of Patagoniaproblematic paleobiological taxonomy |journal=Alcheringa: An Australasian Journal of Systematic Palaeontology |volume=1742 |issue=18 4|pages=1313–1330 516–527|doi=10.1080/1477201903115518.2018.15342881549685 |s2cid=134825554 }}</ref>
* A juvenile mosasaur specimen affected by [[infectious arthritis]] and [[spondyloarthropathy]] is described from the upper [[Maastrichtian]] of [[Antarctica]] by Talevi ''et al.'' (2019), representing the first report of a skeletal pathology of a mosasaur from the [[Southern Hemisphere]].<ref>{{Cite journal|author1=Marianella Talevi |author2=Bruce Rothschild |author3=Marta Fernández |author4=Marcelo Reguero |author5=Matías Mitidieri |year=2019 |title=A pathological scapula in a mosasaur from the upper Maastrichtian of Antarctica: evidence of infectious arthritis and spondyloarthropathy |journal=Cretaceous Research |volume=100 |pages=1–4 |doi=10.1016/j.cretres.2019.03.024 |s2cid=135367471 |url=https://rid.unrn.edu.ar/jspui/handle/20.500.12049/5092 |hdl=11336/125089 |hdl-access=free }}</ref>
* A study on the phylogenetic relationship and evolutionary history of [[caenophidia]]n snakes, as indicated by data from extant taxa and fossil record, is published by Zaher ''et al.'' (2019).<ref>{{Cite journal|author1=Hussam Zaher |author2=Robert W. Murphy |author3=Juan Camilo Arredondo |author4=Roberta Graboski |author5=Paulo Roberto Machado-Filho |author6=Kristin Mahlow |author7=Giovanna G. Montingelli |author8=Ana Bottallo Quadros |author9=Nikolai L. Orlov |author10=Mark Wilkinson |author11=Ya-Ping Zhang |year=2019 |title=Large-scale molecular phylogeny, morphology, divergence-time estimation, and the fossil record of advanced caenophidian snakes (Squamata: Serpentes) |journal=PLoS ONE |volume=14 |issue=5 |pages=e0216148 |doi=10.1371/journal.pone.0216148 |pmid=31075128 |pmc=6512042 }}</ref>
* RevisionA ofstudy [[Dipsadinae|dipsadid]]on snakethe fossilsspatial fromand temporal distribution and evolutionary history of [[GuadeloupeDolichosauridae|dolichosaurs]] Islands is published by BochatonCampbell ''etMekarski, al.''Pierce & Caldwell (2019).<ref>{{Cite journal|author1=CorentinMichelle BochatonCampbell Mekarski |author2=RenaudStephanie BoistelE. Pierce |author3=SandrineMichael GrouardW. |author4=Ivan Ineich |author5=Anne Tresset |author6=Salvador BailonCaldwell |year=2019 |title=FossilSpatiotemporal dipsadiddistributions snakesof fromnon-ophidian theophidiomorphs, Guadeloupewith Islandsimplications (French West-Indies) andfor their interactionsorigin, withradiation, pastand human populationsextinction |journal=GeodiversitasFrontiers in Earth Science |volume=417 |issuearticle-number=12 |pages=501–523245 |doi=10.52523389/geodiversitas2019v41a12feart.2019.00245 |urlbibcode=http://sciencepress2019FrEaS.mnhn.fr/en/periodiques/geodiversitas/41/12.7..245M |doi-access=free }}</ref>
* SnakeA faunastudy fromon the evolution, [[MioceneHomology (biology)|homology]] of the Baikadam and Malyireduction Kalkamanof 1 and 2 localities in northeasternthe [[KazakhstanPelvis|pelvic]], representingelements theand best-documentedhindlimbs Mioceneof snakesnakes assemblageis in Central Asia, will bepublished describedonline by IvanovPalci ''et al.'' (2019).<ref>{{citeCite journal |author1=MartinAlessandro IvanovPalci |author2=DavitMark VasilyanN. Hutchinson |author3=MadelaineMichael BöhmeW. Caldwell |author4=VladimirKrister T. Smith |author5=Michael S. ZazhiginY. Lee |year=2019 |title=MioceneThe snakeshomologies fromand northeasternevolutionary Kazakhstan:reduction newof datathe onpelvis and hindlimbs in snakes, with the evolutionfirst report of snakeossified assemblagespelvic vestiges in Siberiaan anomalepidid (''Liotyphlops beui'') |journal=Historical Biology: An InternationalZoological Journal of Paleobiologythe Linnean Society |volume=in press188 |issue=2 |pages=630–652 1–20|doi=10.10801093/08912963.2018.1446086zoolinnean/zlz098 |doi-access=free }}</ref>
* A study on the evolution of vertebral intercentrum system of snakes, as indicated by data from specimens of ''[[Najash]] rionegrina'' and ''[[Dinilysia]] patagonica'', is published by Garberoglio ''et al.'' (2019).<ref>{{Cite journal|author1=Fernando F. Garberoglio |author2=Raúl O. Gómez |author3=Tiago R. Simões |author4=Michael W. Caldwell |author5=Sebastián Apesteguía |year=2019 |title=The evolution of the axial skeleton intercentrum system in snakes revealed by new data from the Cretaceous snakes ''Dinilysia'' and ''Najash'' |journal=Scientific Reports |volume=9 |issue=1 |article-number=1276 |doi=10.1038/s41598-018-36979-9 |pmid=30718525 |pmc=6362196 |bibcode=2019NatSR...9.1276G }}</ref>
* New specimen of ''Najash rionegrina'', consisting of a partial skull and closely associated vertebrae, is described by Garberoglio ''et al.'' (2019).<ref>{{Cite journal|author1=Fernando F. Garberoglio |author2=Raúl O. Gómez |author3=Sebastián Apesteguía |author4=Michael W. Caldwell |author5=María L. Sánchez |author6=Gonzalo Veiga |year=2019 |title=A new specimen with skull and vertebrae of ''Najash rionegrina'' (Lepidosauria: Ophidia) from the early Late Cretaceous of Patagonia |journal=Journal of Systematic Palaeontology |volume=17 |issue=18 |pages=1533–1550 |doi=10.1080/14772019.2018.1534288 |s2cid=91780191 }}</ref>
* Description of new fossil material (including eight skulls) of ''Najash rionegrina'', and a study on the implications of these fossils for the knowledge of the evolution of the snake body plan, is published by Garberoglio ''et al.'' (2019).<ref>{{Cite journal|author1=Fernando F. Garberoglio |author2=Sebastián Apesteguía |author3=Tiago R. Simões |author4=Alessandro Palci |author5=Raúl O. Gómez |author6=Randall L. Nydam |author7=Hans C. E. Larsson |author8=Michael S. Y. Lee |author9=Michael W. Caldwell |year=2019 |title=New skulls and skeletons of the Cretaceous legged snake ''Najash'', and the evolution of the modern snake body plan |journal=Science Advances |volume=5 |issue=11 |pages=eaax5833 |doi=10.1126/sciadv.aax5833 |pmid=31799393 |pmc=6867888 |bibcode=2019SciA....5.5833G }}</ref>
* A study on the vertebral microanatomy of extinct aquatic snakes belonging to the families [[Nigerophiidae]], [[Palaeophiidae]] and [[Russelophiidae]] is published by Houssaye ''et al.'' (2019).<ref>{{Cite journal|author1=Alexandra Houssaye |author2=Anthony Herrel |author3=Renaud Boistel |author4=Jean-Claude Rage |year=2019 |title=Adaptation of the vertebral inner structure to an aquatic life in snakes: Pachyophiid peculiarities in comparison to extant and extinct forms |journal=Comptes Rendus Palevol |volume=18 |issue=7 |pages=783–799 |doi=10.1016/j.crpv.2019.05.004 |s2cid=209480050 |doi-access=free }}</ref>
* A study on the phylogenetic relationship and evolutionary history of [[caenophidia]]n snakes, as indicated by data from extant taxa and fossil record, is published by Zaher ''et al.'' (2019).<ref>{{Cite journal|author1=Hussam Zaher |author2=Robert W. Murphy |author3=Juan Camilo Arredondo |author4=Roberta Graboski |author5=Paulo Roberto Machado-Filho |author6=Kristin Mahlow |author7=Giovanna G. Montingelli |author8=Ana Bottallo Quadros |author9=Nikolai L. Orlov |author10=Mark Wilkinson |author11=Ya-Ping Zhang |year=2019 |title=Large-scale molecular phylogeny, morphology, divergence-time estimation, and the fossil record of advanced caenophidian snakes (Squamata: Serpentes) |journal=PLOS ONE |volume=14 |issue=5 |pages=e0216148 |doi=10.1371/journal.pone.0216148 |pmid=31075128 |pmc=6512042 |bibcode=2019PLoSO..1416148Z |doi-access=free }}</ref>
* Revision of [[Dipsadinae|dipsadid]] snake fossils from [[Guadeloupe]] Islands is published by Bochaton ''et al.'' (2019).<ref>{{Cite journal|author1=Corentin Bochaton |author2=Renaud Boistel |author3=Sandrine Grouard |author4=Ivan Ineich |author5=Anne Tresset |author6=Salvador Bailon |year=2019 |title=Fossil dipsadid snakes from the Guadeloupe Islands (French West-Indies) and their interactions with past human populations |journal=Geodiversitas |volume=41 |issue=12 |pages=501–523 |doi=10.5252/geodiversitas2019v41a12 |s2cid=198415595 |url=http://sciencepress.mnhn.fr/en/periodiques/geodiversitas/41/12 |doi-access=free }}</ref>
* Snake fauna from the [[Miocene]] of the Baikadam and Malyi Kalkaman 1 and 2 localities in northeastern [[Kazakhstan]], representing the best-documented Miocene snake assemblage in Central Asia, is described by Ivanov ''et al.'' (2019).<ref>{{cite journal |author1=Martin Ivanov |author2=Davit Vasilyan |author3=Madelaine Böhme |author4=Vladimir S. Zazhigin |year=2019 |title=Miocene snakes from northeastern Kazakhstan: new data on the evolution of snake assemblages in Siberia |journal=Historical Biology: An International Journal of Paleobiology |volume=31 |issue=10 |pages=1284–1303 |doi=10.1080/08912963.2018.1446086 |s2cid=90976993 }}</ref>
* Description of fossil snake vertebrae from the late [[Pleistocene]] fissure deposits of the [[Minatogawa Man]] site ([[Okinawa Island]], [[Japan]]) is published by Ikeda ''et al.'' (2019).<ref>{{Cite journal|author1=Tadahiro Ikeda |author2=Akio Takahashi |author3=Makoto Manabe |author4=Yoshikazu Hasegawa |year=2019 |title=Snake vertebrae fossils occurred from the Minatogawa Man site, southern part of Okinawajima Island, Ryukyu Archipelago, southwestern Japan |journal=Bulletin of Gunma Museum of Natural History |volume=23 |pages=21–34 |url=http://www.gmnh.pref.gunma.jp/wp-content/uploads/bulletin23_3.pdf }}</ref>
* Description of Pleistocene snake fossils from the Shanyangzhai Cave ([[Hebei]], [[China]]) will beis published online by Chen ''et al.'' (2019).<ref>{{cite journal |author1=Yu Chen |author2=Yong-Xiang Li |author3=Jing-Song Shi |author4=Yun-Xiang Zhang |author5=Kun Xie |year=2019 |title=Pleistocene fossil snakes (Squamata, Reptilia) from Shanyangzhai Cave, Hebei, China |journal=Historical Biology: An International Journal of Paleobiology |volume=in press33 |issue=5 |pages=699–711 |doi=10.1080/08912963.2019.1658094 |s2cid=202851962 }}</ref>
* RevisionDescription of lizardfossil andmaterial snakeof fossilslizards from the [[Pliocene]]Oligocene siteand Miocene of the [[KanapoiValley of the Lakes]] (central [[KenyaMongolia]]) will beis published by HeadČerňanský & MüllerAugé (2019).<ref>{{cite journal |author1=JasonAndrej J. HeadČerňanský |author2=JohannesMarc Louis MüllerAugé |year=2019 |title=SquamateThe reptilesOligocene fromand Kanapoi:Miocene Faunalfossil evidencelizards for(Reptilia, Squamata) of homininCentral paleoenvironmentsMongolia |journal=Journal of Human EvolutionGeodiversitas |volume=in41 press|issue=24 |pages=811–839 |doi=10.10165252/jgeodiversitas2019v41a24 |s2cid=214190322 |url=http://sciencepress.jhevolmnhn.2018.01.007fr/en/periodiques/geodiversitas/41/24 |pmiddoi-access=29910043free }}</ref>
 
===New taxa===
Line 46 ⟶ 53:
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''[[EgerniaChalcides gillespieaeaugei]]''<ref>{{Citecite journal |author1=Kailah M.Andrej ThornČerňanský |author2=MarkElena NV. HutchinsonSyromyatnikova |author3=MichaelEkaterina ArcherS. Kovalenko |author4=MichaelKonstantin SM. YPodurets |author5=Alexander A. LeeKaloyan |year=20192020 |title=AThe newkey scincidto lizardunderstanding fromthe theEuropean Miocene of''Chalcides'' Northern Australia(Squamata, andScincidae) thecomes evolutionaryfrom Asia: the historylizards of socialthe skinksEast Siberian Tagay locality (Scincidae:Baikal EgerniinaeLake) in Russia |journal=JournalThe of VertebrateAnatomical PaleontologyRecord |volume=39303 |issue=17 |pages=e15778731901–1934 |doi=10.10801002/02724634ar.2019.157787324289 |pmid=31595688 |s2cid=203983366 |doi-access=free }}</ref>
|
Sp. nov
|
Valid
|
Čerňanský ''et al.''
|
Early middle [[Miocene]]
|
|
{{Flag|Russia}}
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A [[skink]], a species of ''[[Chalcides]]''. Announced in 2019; the final version of the article naming it was published in 2020.
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|-
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''[[Egernia gillespieae]]''<ref>{{Cite journal|author1=Kailah M. Thorn |author2=Mark N. Hutchinson |author3=Michael Archer |author4=Michael S. Y. Lee |year=2019 |title=A new scincid lizard from the Miocene of Northern Australia, and the evolutionary history of social skinks (Scincidae: Egerniinae) |journal=Journal of Vertebrate Paleontology |volume=39 |issue=1 |pages=e1577873 |doi=10.1080/02724634.2019.1577873 |s2cid=155763347 }}</ref>
|
Sp. nov
Line 61 ⟶ 85:
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A [[skink]], a species of ''[[Egernia]]''.
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|-
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''[[Eomadtsoia]]''<ref>{{cite journal |author1=Raúl O. Gómez |author2=Fernando F. Garberoglio |author3=Guillermo W. Rougier |year=2019 |title=A new Late Cretaceous snake from Patagonia: Phylogeny and trends in body size evolution of madtsoiid snakes |journal=Comptes Rendus Palevol |volume=18 |issue=7 |pages=771–781 |doi=10.1016/j.crpv.2019.09.003 |hdl=11336/139595 |s2cid=213774502 |hdl-access=free }}</ref>
|
Gen. et sp. nov
|
Valid
|
Gómez, Garberoglio & Rougier
|
[[Late Cretaceous]] ([[Maastrichtian]])
|
[[La Colonia Formation]]
|
{{Flag|Argentina}}
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A [[Madtsoiidae|madtsoiid]] snake. Genus includes new species ''E. ragei''.
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Line 82 ⟶ 124:
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''[[Gurvelus]]''<ref name=PJ531Alifanov>{{Cite journal|author=V. R. Alifanov |year=2019 |title=Lizards of the families Eoxantidae, Ardeosauridae, Globauridae, and Paramacellodidae (Scincomorpha) from the Aptian–Albian of Mongolia |journal=Paleontological Journal |volume=53 |issue=1 |pages=74–88 |doi=10.1134/S0031030119010039 |s2cid=181824832 }}</ref>
|
Gen. et sp. nov
Line 100 ⟶ 142:
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''[[Hoeckosaurus]]''<ref>{{Cite journal|author=Andrej Čerňanský |year=2019 |title=The first potential fossil record of a dibamid reptile (Squamata: Dibamidae): a new taxon from the early Oligocene of Central Mongolia |journal=Zoological Journal of the Linnean Society |volume=Online edition187 |issue=3 |pages=782–799 |doi=10.1093/zoolinnean/zlz047 }}</ref>
|
Gen. et sp. nov
Line 117 ⟶ 159:
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''[[IndrasaurusHongshanxi]]''<ref>{{Citecite journal |author1=JingmaiLiping O’ConnorDong |author2=XiaotingYuan ZhengWang |author3=LipingLijie DongMou |author4=XiaoliGuoze WangZhang |author5=YanSusan WangE. |author6=Xiaomei Zhang |author7=Zhonghe ZhouEvans |year=2019 |title=''Microraptor''A withnew ingestedJurassic lizard suggestsfrom non-specialized digestive functionChina |journal=Current BiologyGeodiversitas |volume=2941 |issue=1416 |pages=2423–2429.e2623–641 |doi=10.10165252/jgeodiversitas2019v41a16 |s2cid=204256127 |url=http://sciencepress.cubmnhn.2019.06.020fr/en/periodiques/geodiversitas/41/16 |pmiddoi-access=31303494free }}</ref>
|
Gen. et sp. nov
Line 123 ⟶ 165:
Valid
|
O’ConnorDong ''et al.''
|
[[Late Jurassic]] ([[Oxfordian (stage)|Oxfordian]])
|
[[Tiaojishan Formation]]
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{{Flag|China}}
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A lizard of uncertain phylogenetic placement. The type species is ''H. xiei''.
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|-
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''[[Indrasaurus]]''<ref>{{Cite journal|author1=Jingmai O'Connor |author2=Xiaoting Zheng |author3=Liping Dong |author4=Xiaoli Wang |author5=Yan Wang |author6=Xiaomei Zhang |author7=Zhonghe Zhou |year=2019 |title=''Microraptor'' with ingested lizard suggests non-specialized digestive function |journal=Current Biology |volume=29 |issue=14 |pages=2423–2429.e2 |doi=10.1016/j.cub.2019.06.020 |pmid=31303494 |s2cid=195887207 |doi-access=free }}</ref>
|
Gen. et sp. nov
|
Valid
|
O'Connor ''et al.''
|
[[Early Cretaceous]]
Line 150 ⟶ 210:
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A lizard belonging to the family [[Globauridae]]. The type species is ''K. aptiana''.
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|-
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''[[Lapparentophis|Lapparentophis ragei]]''<ref>{{Cite journal|author=Romain Vullo |year=2019 |title= A new species of ''Lapparentophis'' from the mid-Cretaceous Kem Kem beds, Morocco, with remarks on the distribution of lapparentophiid snakes |journal=Comptes Rendus Palevol |volume=18 |issue=7 |pages=765–770 |doi=10.1016/j.crpv.2019.08.004 |s2cid=210297438 |doi-access=free }}</ref>
|
Sp. nov
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Valid
|
Vullo
|
[[Cretaceous]] ([[Albian]]–[[Cenomanian]])
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[[Kem Kem Beds]]
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{{Flag|Morocco}}
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An early snake.
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Line 171 ⟶ 249:
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''[[Ophisaurus manchenioi]]''<ref>{{Cite journal|author1=Hugues-Alexandre Blain |author2=Salvador Bailon |year=2019 |title=Extirpation of ''Ophisaurus'' (Anguimorpha, Anguidae) in Western Europe coincided with the disappearance of subtropical ecosystems at the Early-Middle Pleistocene transition |journal=Palaeogeography, Palaeoclimatology, Palaeoecology |volume=520 |pages=96–113 |doi=10.1016/j.palaeo.2019.01.023 |s2cid=135280617 }}</ref>
|
Sp. nov
Line 184 ⟶ 262:
{{Flag|Spain}}
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An [[Anguidae|anguid]] [[lizard]], a species of ''[[Ophisaurus]]''.
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|-
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''[[Palaeopython|Palaeopython helveticus]]''<ref>{{Cite journal |author1=Georgios L. Georgalis |author2=Torsten M. Scheyer |year=2019 |title=A new species of ''Palaeopython'' (Serpentes) and other extinct squamates from the Eocene of Dielsdorf (Zurich, Switzerland) |journal=[[Swiss Journal of Geosciences]] |volume=in112 press|issue=2–3 |pages=383–417 |doi=10.1007/s00015-019-00341-6 |s2cid=195374710 |url=https://www.zora.uzh.ch/id/eprint/171167/24/2019_Georgalis_93.pdf |access-date=2019-08-06 |archive-date=2020-02-28 |archive-url=https://web.archive.org/web/20200228235812/https://www.zora.uzh.ch/id/eprint/171167/24/2019_Georgalis_93.pdf |url-status=dead }}</ref>
|
Sp. nov
|
Valid
In press
|
Georgalis & Scheyer
Line 223 ⟶ 301:
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|
''[[Paradorsetisaurus]]''<ref name=Paradorsetisaurus>{{Cite journal|author=V.R. Alifanov |year=2019 |title=Lizards of the families Dorsetisauridae and Xenosauridae (Anguimorpha) from the Aptian-Albian of Mongolia |journal=Paleontological Journal |volume=53 |issue=2 |pages=183–193 |doi=10.1134/S0031030119020023 |s2cid=195302163 |url=https://elibrary.ru/item.asp?id=37068691 |url-access=subscription }}</ref>
|
Gen. et sp. nov
Line 240 ⟶ 318:
|-
|
''[[Paraxenophis]]''<ref name=Periergophis />
''[[Portunatasaurus]]''<ref>{{Cite journal|author1=Michelle Campbell Mekarski |author2=Dražen Japundžić |author3=Katarina Krizmanić |author4=Michael W. Caldwell |year=2019 |title=Description of a new basal mosasauroid from the Late Cretaceous of Croatia, with comments on the evolution of the mosasauroid forelimb |journal=Journal of Vertebrate Paleontology |volume=39 |issue=1 |pages=e1577872 |doi=10.1080/02724634.2019.1577872 }}</ref>
|
Gen. et sp. nov
|
Valid
|
Georgalis ''et al.''
|
Latest [[Miocene]] or earliest [[Pliocene]]
|
|
{{Flag|Greece}}
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A [[Colubridae|colubrid]] snake. The type species is ''P. spanios''.
|
|-
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''[[Periergophis]]''<ref name=Periergophis>{{cite journal |author1=Georgios L. Georgalis |author2=Andrea Villa |author3=Martin Ivanov |author4=Davit Vasilyan |author5=Massimo Delfino |year=2019 |title=Fossil amphibians and reptiles from the Neogene locality of Maramena (Greece), the most diverse European herpetofauna at the Miocene/Pliocene transition boundary |journal=Palaeontologia Electronica |volume=22 |issue=3 |pages=Article number 22.3.68 |doi=10.26879/908 |s2cid=210621662 |doi-access=free |hdl=2318/1715071 |hdl-access=free }}</ref>
|
Gen. et sp. nov
|
Valid
|
Georgalis ''et al.''
|
Latest [[Miocene]] or earliest [[Pliocene]]
|
|
{{Flag|Greece}}
|
A [[Colubridae|colubrid]] snake. The type species is ''P. micros''.
|
|-
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''[[Portunatasaurus]]''<ref>{{Cite journal|author1=Michelle Campbell Mekarski |author2=Dražen Japundžić |author3=Katarina Krizmanić |author4=Michael W. Caldwell |year=2019 |title=Description of a new basal mosasauroid from the Late Cretaceous of Croatia, with comments on the evolution of the mosasauroid forelimb |journal=Journal of Vertebrate Paleontology |volume=39 |issue=1 |pages=e1577872 |doi=10.1080/02724634.2019.1577872 |s2cid=182237952 }}</ref>
|
Gen. et sp. nov
Line 254 ⟶ 366:
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An [[aigialosaur]] [[mosasaur]]oid. Genus includes new species ''P. krambergeri''.
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|-
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''[[Sardophis]]''<ref>{{Cite journal|author1=Georgios L. Georgalis |author2=Marisa Arca |author3=Lorenzo Rook |author4=Caterinella Tuveri |author5=Massimo Delfino |year=2019 |title=A new colubroid snake (Serpentes) from the early Pleistocene of Sardinia, Italy |journal=Bollettino della Società Paleontologica Italiana |volume=58 |issue=3 |pages=277–294 |doi=10.4435/BSPI.2019.19 |doi-broken-date=4 July 2025 }}</ref>
|
Gen. et sp. nov
|
Valid
|
Georgalis & Delfino ''in'' Georgalis ''et al.''
|
Early [[Pleistocene]]
|
|
{{Flag|Italy}}
|
A snake, a member of [[Colubroidea]] of uncertain phylogenetic placement. The type species is ''S. elaphoides''.
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|-
Line 276 ⟶ 405:
 
==Ichthyosauromorphs==
 
* Two new specimens of ''[[Eretmorhipis]] carrolldongi'', revealing superficial [[Convergent evolution|convergence]] with the modern [[platypus]], are described from the [[Lower Triassic]] [[Jialingjiang Formation]] ([[China]]) by Cheng ''et al.'' (2019).<ref>{{cite journal |author1=Long Cheng |author2=Ryosuke Motani |author3=Da-yong Jiang |author4=Chun-bo Yan |author5=Andrea Tintori |author6=Olivier Rieppel |year=2019 |title=Early Triassic marine reptile representing the oldest record of unusually small eyes in reptiles indicating non-visual prey detection |journal=Scientific Reports |volume=9 |issue=1 |pages=Article number 152 |doi=10.1038/s41598-018-37754-6 |pmid=30679783 |pmc=6345829 }}</ref>
===Research===
* A study on the phylogenetic relationships of [[ichthyosaur]]s will be published by Moon (2019).<ref>{{cite journal |author=Benjamin C. Moon |year=2019 |title=A new phylogeny of ichthyosaurs (Reptilia: Diapsida) |journal=Journal of Systematic Palaeontology |volume=17 |issue=2 |pages=129–155 |doi=10.1080/14772019.2017.1394922 |hdl=1983/463e9f78-10b7-4262-9643-0454b4aa7763 }}</ref>
* ATwo studynew onspecimens theof evolution''[[Eretmorhipis]] ofcarrolldongi'', ichthyosaurrevealing bodysuperficial forms[[Convergent andevolution|convergence]] onwith itsthe impactmodern on[[platypus]], theare energydescribed demandsfrom ofthe ichthyosaur[[Lower swimmingTriassic]] is[[Jialingjiang publishedFormation]] ([[China]]) by GutarraCheng ''et al.'' (2019).<ref>{{Citecite journal |author1=SusanaLong GutarraCheng |author2=BenjaminRyosuke C. MoonMotani |author3=Imran A.Da-yong RahmanJiang |author4=ColinChun-bo PalmerYan |author5=StephanAndrea LautenschlagerTintori |author6=AlisonOlivier J. Brimacombe |author7=Michael J. BentonRieppel |year=2019 |title=EffectsEarly ofTriassic bodymarine planreptile evolution onrepresenting the hydrodynamicoldest dragrecord andof energyunusually requirementssmall of swimmingeyes in ichthyosaursreptiles |journal=Proceedingsindicating ofnon-visual theprey Royaldetection Society|journal=Scientific B: Biological SciencesReports |volume=2869 |issue=18981 |pagesarticle-number=Article ID 20182786152 |doi=10.10981038/rspb.2018.2786s41598-018-37754-6 |pmid=3083686730679783 |pmc=64583256345829 |bibcode=2019NatSR...9..152C }}</ref>
* A study on the flexibilityphylogenetic and functionrelationships of [[Ichthyosauria|ichthyosaur]]s tails,will as indicated by comparisons with shark tails, isbe published by Crofts, Shehata & FlammangMoon (2019).<ref>{{Citecite journal|author1=S. B. Crofts |author2author=R.Benjamin Shehata |author3=BC. FlammangMoon |year=2019 |title=FlexibilityA ofnew heterocercal tails: what can the functional morphologyphylogeny of sharkichthyosaurs tails(Reptilia: tell us about ichthyosaur swimming?Diapsida) |journal=IntegrativeJournal of OrganismalSystematic BiologyPalaeontology |volume=117 |issue=12 |pages=obz002129–155 |doi=10.10931080/iob14772019.2017.1394922 |hdl=1983/obz002463e9f78-10b7-4262-9643-0454b4aa7763 |s2cid=90912678 |url=https://research-information.bris.ac.uk/ws/files/142491822/Typescript_A_new_phylogeny_of_ichthyosaurs.pdf }}</ref>
* A study on the effectsevolution of methodology,ichthyosaur missingbody dataforms and exceptionalon preservationits of fossil specimens in [[lagerstätte]]nimpact on knownthe morphologicalenergy diversitydemands of fossil animals, asichthyosaur indicated by fossil record of ichthyosaurs,swimming is published by Flannery SutherlandGutarra ''et al.'' (2019).<ref>{{Cite journal|author1=JosephSusana T. Flannery SutherlandGutarra |author2=Benjamin C. Moon |author3=ThomasImran LA. StubbsRahman |author4=Colin Palmer |author5=Stephan Lautenschlager |author6=Alison J. Brimacombe |author7=Michael J. Benton |year=2019 |title=DoesEffects exceptionalof preservationbody distortplan ourevolution viewon ofthe disparityhydrodynamic indrag theand fossilenergy record?requirements of swimming in ichthyosaurs |journal=Proceedings of the Royal Society B: Biological Sciences |volume=286 |issue=18971898 |pagesarticle-number=Article ID 2019009120182786 |doi=10.1098/rspb.20192018.00912786 |pmid=3096385030836867 |pmc=64089026458325 }}</ref>
* A study on athe putativeflexibility Cretaceousand [[mosasaur]]function ''"[[Globidens]]"of timorensis''ichthyosaur fromtails, [[Timor]]as is publishedindicated by Muldercomparisons &with Jagtshark (2019)tails, whois considerpublished thisby taxon to be of [[Triassic]] ageCrofts, andShehata reinterpret& itFlammang as an ichthyosaur with affinities to the genera ''[[Tholodus]]'' and ''[[Xinminosaurus]]''(2019).<ref>{{Cite journal|author1=EricS. W.AB. MulderCrofts |author2=John WR.M Shehata |author3=B. JagtFlammang |year=2019 |title=''Globidens(?)Flexibility timorensis''of E.heterocercal VONtails: HUENE,what 1935:can notthe afunctional durophagousmorphology mosasaur,of butshark antails enigmatictell Triassicus about ichthyosaur swimming? |journal=NeuesIntegrative JahrbuchOrganismal für Geologie und Paläontologie - AbhandlungenBiology |volume=2931 |issue=1 |pages=107–116obz002 |doi=10.11271093/njgpaiob/2019/0835obz002 |pmid=33791519 |pmc=7671117 }}</ref>
* A study on the prevalenceeffects of methodology, missing data and distributionexceptional preservation of pathologiesfossil specimens in the[[lagerstätte]]n skeletonson known morphological diversity of ichthyosaursfossil fromanimals, theas Lowerindicated Jurassicby [[Posidonienschieferfossil Formation]]record ([[Germany]])of ichthyosaurs, is published by Pardo-Pérez,Flannery KearSutherland &''et Maxwellal.'' (2019).<ref>{{Cite journal|author1=JudithJoseph MT. Pardo-PérezFlannery Sutherland |author2=Benjamin KearC. Moon |author3=ErinThomas L. Stubbs |author4=Michael EJ. MaxwellBenton |year=2019 |title=PalaeoepidemiologyDoes inexceptional extinctpreservation vertebratedistort populations:our factorsview influencingof skeletal healthdisparity in Jurassicthe marinefossil reptilesrecord? |journal=Proceedings of the Royal Society OpenB: Biological ScienceSciences |volume=6286 |issue=71897 |pagesarticle-number=Article ID 19026420190091 |doi=10.1098/rsosrspb.2019.1902640091 |pmid=3141773230963850 |pmc=66895716408902 }}</ref>
* SecondA specimenstudy ofon a putative Cretaceous [[mosasaur]] ''"[[WahlisaurusGlobidens]]" massaraetimorensis'' is reported from a quarry in [[SomersetTimor]] ([[Unitedis Kingdom]]published by Mulder & Jagt (2019), fromwho theconsider basethis oftaxon theto [[Bluebe Lias]]of Formation ([[Triassic]]–[[Jurassic]] boundary) by Lomaxage, Evansand &reinterpret Carpenterit (2019),as extendingan knownichthyosaur geographicwith andaffinities stratigraphicto rangethe ofgenera the''[[Tholodus]]'' speciesand ''[[Xinminosaurus]]''.<ref>{{citeCite journal |author1=DeanEric RW.A. LomaxMulder |author2=MarkJohn EvansW.M. |author3=Simon CarpenterJagt |year=2019 |title=An''Globidens(?) ichthyosaurtimorensis'' fromE. theVON UK Triassic–JurassicHUENE, boundary1935: Anot seconda specimendurophagous ofmosasaur, thebut leptonectidan ichthyosaurenigmatic ''WahlisaurusTriassic massarae'' Lomax 2016ichthyosaur |journal=GeologicalNeues JournalJahrbuch für Geologie und Paläontologie - Abhandlungen |volume=54293 |issue=1 |pages=83–90107–116 |doi=10.10021127/gj.3155njgpa/2019/0835 |s2cid=199112423 }}</ref>
* PartialA skeletonstudy on the prevalence and distribution of apathologies largein the skeletons of ichthyosaurichthyosaurs from the [[Lower Jurassic]] ([[Sinemurian]])Posidonienschiefer of [[WarwickshireFormation]], ([[EnglandGermany]] is described) by LomaxPardo-Pérez, PorroKear & LarkinMaxwell (2019), who assign this specimen to the species ''[[Protoichthyosaurus]] prostaxalis''.<ref>{{citeCite journal |author1=DeanJudith RM. LomaxPardo-Pérez |author2=LauraBenjamin B. PorroKear |author3=NigelErin RE. LarkinMaxwell |year=2019 |title=DescriptivePalaeoepidemiology anatomyin ofextinct thevertebrate largestpopulations: knownfactors specimeninfluencing ofskeletal ''Protoichthyosaurushealth prostaxalis''in (Reptilia:Jurassic Ichthyosauria)marine includingreptiles computed|journal=Royal tomographySociety andOpen digital reconstruction of a three-dimensional skullScience |journalvolume=PeerJ6 |volumeissue=7 |pagesarticle-number=e6112190264 |doi=10.77171098/peerjrsos.6112190264 |pmid=3064369031417732 |pmc=63293386689571 |bibcode=2019RSOS....690264P }}</ref>
* A [[neonate]]Second specimen of ''[[IchthyosaurusWahlisaurus]] communismassarae'' is describedreported from a quarry in [[Somerset]] ([[United Kingdom]]), from the base of the [[Blue Lias]] Formation ([[Triassic]]–[[Jurassic]] boundary) by Lomax, ''etEvans al.''& Carpenter (2019), extending known geographic and stratigraphic range of the species.<ref>{{cite journal |author1=Dean R. Lomax |author2=NigelMark R. LarkinEvans |author3=Ian Boomer |author4=Steven Dey |author5=PhilipSimon CopestakeCarpenter |year=2019 |title=TheAn firstichthyosaur knownfrom neonatethe ''IchthyosaurusUK communis''Triassic–Jurassic skeletonboundary: aA rediscoveredsecond specimen fromof the Lowerleptonectid Jurassic,ichthyosaur UK''Wahlisaurus |journal=Historicalmassarae'' Biology:Lomax An2016 International|journal=Geological Journal of Paleobiology |volume=3154 |issue=51 |pages=600–60983–90 |doi=10.10801002/08912963gj.2017.13824883155 |urls2cid=http://pure134448865 |doi-oai.bham.ac.uk/ws/files/43380733/Lomax_etal_2017_HistBiol_I.communis_Neonate_Figs.pdfaccess=free }}</ref>
* Partial skeleton of a large ichthyosaur from the [[Lower Jurassic]] ([[Sinemurian]]) of [[Warwickshire]], [[England]] is described by Lomax, Porro & Larkin (2019), who assign this specimen to the species ''[[Protoichthyosaurus]] prostaxalis''.<ref>{{cite journal |author1=Dean R. Lomax |author2=Laura B. Porro |author3=Nigel R. Larkin |year=2019 |title=Descriptive anatomy of the largest known specimen of ''Protoichthyosaurus prostaxalis'' (Reptilia: Ichthyosauria) including computed tomography and digital reconstruction of a three-dimensional skull |journal=PeerJ |volume=7 |article-number=e6112 |doi=10.7717/peerj.6112 |pmid=30643690 |pmc=6329338 |doi-access=free }}</ref>
* A study on the variation of the hindfin [[Morphology (biology)|morphology]] in the specimens of ''Ichthyosaurus'' and on its taxonomic utility is published by Massare & Lomax (2019).<ref>{{cite journal |author1=Judy A. Massare |author2=Dean R. Lomax |year=2019 |title=Hindfins of ''Ichthyosaurus'': effects of large sample size on 'distinct' morphological characters |journal=Geological Magazine |volume=156 |issue=4 |pages=725–744 |doi=10.1017/S0016756818000146 |url=https://www.research.manchester.ac.uk/portal/en/publications/hindfins-of-ichthyosaurus(c6032eb0-a39b-4f48-8c1b-eb2f81c23d1f).html }}</ref>
* A study on the bone microstructureanatomy of the skeletonskull of a specimenroof of ''[[Stenopterygius]]Protoichthyosaurus quadriscissusprostaxalis'', fromincorporating thedata Lowerfrom Jurassica [[Posidoniapreviously Shale]]unrecognized ([[Germany]])specimen, is published online by AndersonLomax, ''etMassare al.''& Evans (2019).<ref>{{Citecite journal |author1=KatherineDean LR. AndersonLomax |author2=PatrickJudy SA. DruckenmillerMassare |author3=GregoryMark M. Erickson |author4=Erin E. MaxwellEvans |year=2019 |title=SkeletalNew microstructureinformation on the skull roof of ''Stenopterygius quadriscissusProtoichthyosaurus'' (Reptilia,: Ichthyosauria) fromand theintraspecific Posidonienschiefervariation (Posidoniain Shale,some Lowerdermal Jurassic)skull of Germanyelements |journal=PalaeontologyGeological Magazine |volume=62157 |issue=34 |pages=433–449640–650 |doi=10.11111017/pala.12408S0016756819001225 |s2cid=210267260 }}</ref>
* A study on the anatomy of an [[Ophthalmosauridae|ophthalmosauridneonate]] rostrumspecimen fragment from theof ''[[Upper JurassicIchthyosaurus]] ([[Oxfordiancommunis'' (stage)|Oxfordian]])is in the Morawica quarry in the [[Świętokrzyskie Mountains]] ([[Poland]]), and on its implications for reconstructing the internal [[Morphology (biology)|morphology]] of the ophthalmosaurid cranial region and inferring the functional adaptations and palaeoecology of these reptiles, will be publisheddescribed by Tyborowski,Lomax Skrzycki''et & Decal.'' (2019).<ref>{{cite journal |author1=DanielDean TyborowskiR. Lomax |author2=PiotrNigel R. SkrzyckiLarkin |author3=MarekIan DecBoomer |author4=Steven Dey |author5=Philip Copestake |year=2019 |title=InternalThe structurefirst ofknown ichthyosaurneonate rostrum''Ichthyosaurus fromcommunis'' theskeleton: Uppera Jurassicrediscovered ofspecimen Polandfrom withthe commentsLower on ecomorphological adaptations of ophthalmosauridJurassic, skullUK |journal=Historical Biology: An International Journal of Paleobiology |volume=in press31 |issue=5 |pages=600–609 1–10|doi=10.1080/08912963.20182017.1382488 |s2cid=89922569 |url=http://pure-oai.bham.ac.uk/ws/files/43380733/Lomax_etal_2017_HistBiol_I.communis_Neonate_Figs.1559308pdf }}</ref>
* A revisionstudy on the variation of the hindfin [[typeMorphology series(biology)|morphology]] ofin all threethe speciesspecimens of ''[[Undorosaurus]]Ichthyosaurus'' and on its taxonomic utility is published by ZverkovMassare & EfimovLomax (2019).<ref>{{cite journal |author1=NikolayJudy GA. ZverkovMassare |author2=VladimirDean MR. EfimovLomax |year=2019 |title=RevisionHindfins of ''UndorosaurusIchthyosaurus'',: aeffects mysteriousof Latelarge Jurassicsample ichthyosaursize ofon the'distinct' Borealmorphological Realmcharacters |journal=Journal of SystematicGeological PalaeontologyMagazine |volume=17156 |issue=144 |pages=963–993725–744 |doi=10.10801017/14772019S0016756818000146 |bibcode=2019GeoM.2018.1515793156..725M |s2cid=134945422 |url=https://figsharewww.comresearch.manchester.ac.uk/articlesportal/Revision_of_i_Undorosaurus_i_a_mysterious_Late_Jurassic_ichthyosaur_of_the_Boreal_Realmen/7624868publications/hindfins-of-ichthyosaurus(c6032eb0-a39b-4f48-8c1b-eb2f81c23d1f).html }}</ref>
* A study on the taxonomybone and phylogenymicrostructure of ichthyosaursthe belongingskeleton toof thea genusspecimen of ''[[ArthropterygiusStenopterygius]] quadriscissus'' from the Lower Jurassic [[Posidonia Shale]] ([[Germany]]) is published by ZverkovAnderson &''et Prilepskayaal.'' (2019).<ref>{{citeCite journal |author1=NikolayKatherine GL. ZverkovAnderson |author2=NatalyaPatrick S. Druckenmiller |author3=Gregory M. Erickson |author4=Erin E. PrilepskayaMaxwell |year=2019 |title=ASkeletal prevalencemicrostructure of ''ArthropterygiusStenopterygius quadriscissus'' (Ichthyosauria:Reptilia, OphthalmosauridaeIchthyosauria) infrom the LatePosidonienschiefer Jurassic—earliest(Posidonia CretaceousShale, ofLower theJurassic) Borealof RealmGermany |journal=PeerJPalaeontology |volume=762 |issue=3 |pages=e6799433–449 |doi=10.77171111/peerjpala.6799 |pmid=3110605212408 |pmcs2cid=6497043135229818 }}</ref>
* NewA fossilstudy remainson ofthe ''[[PlatypterygiusOntogeny|ontogenetic]] sachicarum''variation (ain newthe skullanatomy andof associatedthe postcranial remainsbraincases of uppermembers [[Barremian]]of age)the aregenus described''Stenopterygius'' fromis Villapublished deby Leyva,Miedema [[Colombia]] by& Maxwell ''et al.'' (2019), representing the first documented postcranial remains of this species.<ref>{{Citecite journal |author1=ErinFeiko E. MaxwellMiedema |author2=DirleyErin CortésE. |author3=Pedro Patarroyo |author4=Mary Luz Parra RugeMaxwell |year=2019 |title=AOntogeny newof specimenthe ofbraincase in ''Platypterygius sachicarumStenopterygius'' (Reptilia, Ichthyosauria) from the EarlyLower CretaceousJurassic of Colombia and its phylogenetic implicationsGermany |journal=Journal of Vertebrate Paleontology |volume=39 |issue=14 |pages=e1577875e1675164 |doi=10.1080/02724634.2019.15778751675164 |s2cid=209439995 |doi-access=free }}</ref>
* Description of new ophthalmosaurid fossils from the Upper Jurassic of the Slottsmøya Member Lagerstätte ([[Spitsbergen]], [[Norway]]) and a study on the phylogenetic relationships of ophthalmosaurid specimens from the Slottsmøya Member Lagerstätte is published by Delsett ''et al.'' (2019).<ref>{{cite journal |author1=Lene L. Delsett |author2=Aubrey J. Roberts |author3=Patrick S. Druckenmiller |author4=Jørn H. Hurum |year=2019 |title=Osteology and phylogeny of Late Jurassic ichthyosaurs from the Slottsmøya Member Lagerstätte (Spitsbergen, Svalbard) |journal=Acta Palaeontologica Polonica |volume=64 |issue=4 |pages=717–743 |doi=10.4202/app.00571.2018 |s2cid=210294877 |doi-access=free |hdl=10852/76089 |hdl-access=free }}</ref>
* A revision of the [[type series]] of all three species of ''[[Undorosaurus]]'' is published by Zverkov & Efimov (2019).<ref>{{cite journal |author1=Nikolay G. Zverkov |author2=Vladimir M. Efimov |year=2019 |title=Revision of ''Undorosaurus'', a mysterious Late Jurassic ichthyosaur of the Boreal Realm |journal=Journal of Systematic Palaeontology |volume=17 |issue=14 |pages=1183–1213 |doi=10.1080/14772019.2018.1515793 |bibcode=2019JSPal..17.1183Z |s2cid=91912834 |url=https://figshare.com/articles/Revision_of_i_Undorosaurus_i_a_mysterious_Late_Jurassic_ichthyosaur_of_the_Boreal_Realm/7624868 }}</ref>
* A study on the taxonomy and phylogeny of ichthyosaurs belonging to the genus ''[[Arthropterygius]]'' is published by Zverkov & Prilepskaya (2019).<ref>{{cite journal |author1=Nikolay G. Zverkov |author2=Natalya E. Prilepskaya |year=2019 |title=A prevalence of ''Arthropterygius'' (Ichthyosauria: Ophthalmosauridae) in the Late Jurassic—earliest Cretaceous of the Boreal Realm |journal=PeerJ |volume=7 |article-number=e6799 |doi=10.7717/peerj.6799 |pmid=31106052 |pmc=6497043 |doi-access=free }}</ref>
* New fossil remains of ''[[Platypterygius]] sachicarum'' (a new skull and associated postcranial remains of upper [[Barremian]] age) are described from Villa de Leyva, [[Colombia]] by Maxwell ''et al.'' (2019), representing the first documented postcranial remains of this species.<ref>{{Cite journal|author1=Erin E. Maxwell |author2=Dirley Cortés |author3=Pedro Patarroyo |author4=Mary Luz Parra Ruge |year=2019 |title=A new specimen of ''Platypterygius sachicarum'' (Reptilia, Ichthyosauria) from the Early Cretaceous of Colombia and its phylogenetic implications |journal=Journal of Vertebrate Paleontology |volume=39 |issue=1 |pages=e1577875 |doi=10.1080/02724634.2019.1577875 |s2cid=146059015 }}</ref>
 
===New taxa===
{| class="wikitable sortable" align="center" width="100%"
|-
! Name
! Novelty
! Status
! Authors
! Age
! Type locality
! Country
! Notes
! Images
|-
|
''[[Arthropterygius|Arthropterygius thalassonotus]]''<ref>{{cite journal |author1=Lisandro Campos |author2=Marta S. Fernández |author3=Yanina Herrera |year=2020 |title=A new ichthyosaur from the Late Jurassic of north-west Patagonia (Argentina) and its significance for the evolution of the narial complex of the ophthalmosaurids |journal=Zoological Journal of the Linnean Society |volume=188 |issue=1 |pages=180–201 |doi=10.1093/zoolinnean/zlz095 |doi-access=free }}</ref>
|
Sp. nov
|
Valid
|
Campos, Fernández & Herrera
|
[[Late Jurassic]]
|
[[Vaca Muerta]] Formation
|
{{Flag|Argentina}}
|
|
|-
|
''[[Chaohusaurus|Chaohusaurus brevifemoralis]]''<ref>{{cite journal |author1=Jian-dong Huang |author2=Ryosuke Motani |author3=Da-yong Jiang |author4=Andrea Tintori |author5=Olivier Rieppel |author6=Min Zhou |author7=Xin-Xin Ren |author8=Rong Zhang |year=2019 |title=The new ichthyosauriform ''Chaohusaurus brevifemoralis'' (Reptilia, Ichthyosauromorpha) from Majiashan, Chaohu, Anhui Province, China |journal=PeerJ |volume=7 |article-number=e7561 |doi=10.7717/peerj.7561 |pmid=31565558 |pmc=6741286 |doi-access=free }}</ref>
|
Sp. nov
|
Valid
|
Huang ''et al.''
|
[[Early Triassic]]
|
|
{{Flag|China}}
|
|[[File:Chaohusaurus_brevifemoralis_Holotype_NGMC.jpg|center|frameless]]
|-
|}
 
==Sauropterygians==
===Research===
* A study on the microstructurebone [[histology]] of ribssauropterygians, and vertebraeon its implications for the knowledge of [[Middlethe Triassic]]evolution sauropterygiansof willdiving beadaptations publishedof bymembers Kleinof this group, Canovilleis &published Houssayeby Fleischle ''et al.'' (2019).<ref>{{Cite journal|author1=NicoleCorinna KleinV. Fleischle |author2=AuroreP. CanovilleMartin Sander |author3=AlexandraTanja Wintrich |author4=Kai R. HoussayeCaspar |year=2019 |title=MicrostructureHematological ofconvergence vertebrae,between ribs,Mesozoic andmarine gastraliareptiles of(Sauropterygia) Triassicand sauropterygiansextant aquatic Newamniotes insightselucidates intodiving the microanatomical processes involvedadaptations in aquatic adaptations of marine reptilesplesiosaurs |journal=The Anatomical RecordPeerJ |volume=in press7 |issue= |pagesarticle-number=e8022 |doi=10.10027717/arpeerj.241408022 |pmid=3098982831763069 |pmc=6873879 |doi-access=free }}</ref>
* A study on the [[taphonomy]]microstructure of sauropterygianribs specimensand fromvertebrae theof [[Middle Triassic fossil deposit of Winterswijk (the [[Netherlands]]) sauropterygians is published by HeijneKlein, KleinCanoville & SanderHoussaye (2019).<ref>{{Cite journal|author1=JelleNicole HeijneKlein |author2=NicoleAurore KleinCanoville |author3=P.Alexandra Martin SanderHoussaye |year=2019 |title=TheMicrostructure uniquelyof diversevertebrae, taphonomyribs, and gastralia of theTriassic marinesauropterygians reptile skeletonsNew (Sauropterygia)insights frominto the Lowermicroanatomical Muschelkalkprocesses (Anisian)involved ofin Winterswijk,aquatic Theadaptations Netherlandsof marine reptiles |journal=PalZThe Anatomical Record |volume=93302 |issue=110 |pages=69–921770–1791 |doi=10.10071002/s12542ar.24140 |pmid=30989828 |s2cid=117721335 |doi-018-0438-0access=free }}</ref>
* A study on the [[taphonomy]] of sauropterygian specimens from the Middle Triassic fossil deposit of Winterswijk (the [[Netherlands]]) is published by Heijne, Klein & Sander (2019).<ref>{{Cite journal|author1=Jelle Heijne |author2=Nicole Klein |author3=P. Martin Sander |year=2019 |title=The uniquely diverse taphonomy of the marine reptile skeletons (Sauropterygia) from the Lower Muschelkalk (Anisian) of Winterswijk, The Netherlands |journal=PalZ |volume=93 |issue=1 |pages=69–92 |doi=10.1007/s12542-018-0438-0 |s2cid=133992814 }}</ref>
* The first subadult specimen of ''[[Psephochelys]] polyosteoderma'', representing the most complete specimen of this taxon reported so far and providing new information on the anatomy of this taxon, is described from south-western [[China]] by Wang, Ma & Li (2019).<ref>{{cite journal |author1=Wei Wang |author2=Feimin Ma |author3=Chun Li |year=2019 |title=First subadult specimen of ''Psephochelys polyosteoderma'' (Sauropterygia, Placodontia) implies turtle-like fusion pattern of the carapace |journal=Papers in Palaeontology |volume=6 |issue=2 |pages=251–264 |doi=10.1002/spp2.1293 |s2cid=212865203 }}</ref>
* The first adult specimen of ''[[Sinocyamodus]] xinpuensis'' reported so far is described by Wang, Li & Wu (2019).<ref>{{cite journal |author1=Wei Wang |author2=Chun Li |author3=Xiao-Chun Wu |year=2019 |title=An adult specimen of ''Sinocyamodus xinpuensis'' (Sauropterygia: Placodontia) from Guanling, Guizhou, China |journal=Zoological Journal of the Linnean Society |volume=185 |issue=3 |pages=910–924 |doi=10.1093/zoolinnean/zly080 }}</ref>
* Two [[pachypleurosaur]] specimens are described from the Lashio Basin ([[Myanmar]]) by San ''et al.'' (2019), representing the first [[Triassic]] vertebrate fossils from Myanmar reported so far.<ref>{{cite journal |author1=Khaing Khaing San |author2=Nicholas C. Fraser |author3=Davide Foffa |author4=Olivier Rieppel |author5=Stephen L. Brusatte |year=2019 |title=The first Triassic vertebrate fossils from Myanmar: Pachypleurosaurs in a marine limestone |journal=Acta Palaeontologica Polonica |volume=64 |issue=2 |pages=357–362 |doi=10.4202/app.00594.2019 |s2cid=149583826 |doi-access=free |hdl=20.500.11820/5fa5756a-4e76-4f62-b444-413ff379fbbd |hdl-access=free }}</ref>
* A studylarge onmarine lifesauropterygian historybelonging ofor related to ''[[NothosaurusNothosaur]]'',oidea asis indicateddescribed byfrom growththe curvesTriassic determined(probably from[[Olenekian]]) [[Humerus|humeralSulphur Mountain Formation]] ([[histologyBritish Columbia]], and[[Canada]]) onby itsScheyer, implicationsNeuman for& inferringBrinkman reproduction mode(2019), representing one of thisthe animal,oldest isrecords publishedof bySauropterygia Griebelerand &the Kleinnorthernmost (2019)occurrence of such animals in the Triassic.<ref>{{Citecite journal |author1=EvaTorsten M. GriebelerScheyer |author2=NicoleAndrew KleinG. Neuman |author3=Donald B. Brinkman |year=2019 |title=Life‐historyA strategieslarge indicatemarine live‐bearingeosauropterygian in ''Nothosaurus''reptile (Sauropterygia)with affinities to nothosauroid diapsids from the Early Triassic of British Columbia, Canada |journal=PalaeontologyActa Palaeontologica Polonica |volume=6264 |issue=4 |pages=697–713745–755 |doi=10.11114202/palaapp.1242500599.2019 |s2cid=204794697 |doi-access=free }}</ref>
* DescriptionA ofstudy microbodieson extractedlife from a bonehistory of ''[[Nothosaurus]]'', fromas theindicated by growth curves determined from [[Middle TriassicHumerus|humeral]] of [[Polandhistology]], reportedand ason morphologicallyits consistentimplications withfor boneinferring cellsreproduction mode of present-daythis vertebratesanimal, will beis published by SurmikGriebeler ''et& al.''Klein (2019).<ref>{{Cite journal|author1=D.Eva Surmik |author2=M. DulskiGriebeler |author3author2=B.Nicole Kremer |author4=J. Szade |author5=R. PawlickiKlein |year=2019 |title=IronLife-mediatedhistory deep-timestrategies preservationindicate of osteocyteslive-bearing in a''Nothosaurus'' Middle Triassic reptile bone(Sauropterygia) |journal=Historical Biology: An International Journal of PaleobiologyPalaeontology |volume=in press62 |issue=4 |pages=697–713 1–8|doi=10.10801111/08912963pala.2019.159988412425 |s2cid=133629062 }}</ref>
* AnDescription articulatedof juvenilemicrobodies specimenextracted from a bone of ''[[YunguisaurusNothosaurus]] liae'', providingfrom newthe information[[Middle onTriassic]] theof anatomy[[Poland]], ofreported thisas species,morphologically isconsistent describedwith frombone thecells [[Ladinian]]of [[Falangpresent-day Formation]]vertebrates, ([[China]])is published online by WangSurmik ''et al.'' (2019).<ref>{{citeCite journal |author1=XueD. WangSurmik |author2=HaoM. LuDulski |author3=Da-YongB. JiangKremer |author4=MinJ. ZhouSzade |author5=Zuo-YuR. SunPawlicki |year=2019 |title=AIron-mediated newdeep-time specimenpreservation of ''Yunguisaurus''osteocytes (Reptilia;in Sauropterygia)a from the Ladinian (Middle Triassic) Zhuganporeptile Member,bone Falang|journal=Historical Formation,Biology: Guizhou,An ChinaInternational and the restudyJournal of ''Dingxiaosaurus'' |journal=PalaeoworldPaleobiology |volume=2833 |issue=2 |pages=186–193 |doi=10.10161080/j.palwor08912963.2019.05.0061599884 |s2cid=146090823 }}</ref>
* AAn studyarticulated onjuvenile specimen of ''[[hydrodynamicsYunguisaurus]] ofliae'', neckproviding lengthnew andinformation thicknesson inthe plesiosaursanatomy of this species, is publisheddescribed from the [[Ladinian]] [[Falang Formation]] ([[China]]) by TroelsenWang ''et al.'' (2019).<ref>{{Citecite journal |author1=PernilleXue V. TroelsenWang |author2=David M.Hao WilkinsonLu |author3=MehdiDa-Yong SeddighiJiang |author4=DavidMin R. AllansonZhou |author5=Peter L.Zuo-Yu FalkinghamSun |year=2019 |title=FunctionalA morphologynew and hydrodynamicsspecimen of plesiosaur''Yunguisaurus'' necks:(Reptilia; DoesSauropterygia) sizefrom matter?the |journal=JournalLadinian (Middle Triassic) Zhuganpo Member, Falang Formation, Guizhou, China and the restudy of Vertebrate''Dingxiaosaurus'' Paleontology|journal=Palaeoworld |volume=3929 |issue=21 |pages=e1594850137–150 |doi=10.10801016/02724634j.palwor.2019.159485005.006 |s2cid=181711576 }}</ref>
* A study on [[hydrodynamics]] of neck length and thickness in plesiosaurs is published by Troelsen ''et al.'' (2019).<ref>{{Cite journal|author1=Pernille V. Troelsen |author2=David M. Wilkinson |author3=Mehdi Seddighi |author4=David R. Allanson |author5=Peter L. Falkingham |year=2019 |title=Functional morphology and hydrodynamics of plesiosaur necks: Does size matter? |journal=Journal of Vertebrate Paleontology |volume=39 |issue=2 |pages=e1594850 |doi=10.1080/02724634.2019.1594850 |s2cid=181587237 |url=https://researchonline.ljmu.ac.uk/id/eprint/10242/1/Troelsen%20et%20al.pdf }}</ref>
* Pathological fusions of neck vertebrae are reported in four plesiosaur specimens from different geological horizons by Sassoon (2019).<ref>{{cite journal |author=Judyth Sassoon |year=2019 |title=Congenital and late onset vertebral fusions in long necked plesiosaurs: The first report of spondylosis deformans in Sauropterygians |journal=Palaeontologia Electronica |volume=22 |issue=1 |pages=Article number 22.1.1 |doi=10.26879/913 }}</ref>
* FourPathological teethfusions of aneck freshwater plesiosaurvertebrae are describedreported fromin thefour [[Middleplesiosaur Jurassic]]specimens [[Xinhefrom Formation]]different ([[Gansu]],geological [[China]])horizons by Gao ''et al.''Sassoon (2019).<ref>{{cite journal |author1author=TingJudyth Gao |author2=Da-Qing Li |author3=Long-Feng Li |author4=Jing-Tao YangSassoon |year=2019 |title=TheCongenital firstand recordlate ofonset freshwatervertebral plesiosaurianfusions fromin thelong Middlenecked Jurassicplesiosaurs: ofThe Gansu,first NWreport China,of withspondylosis itsdeformans implicationsin to the local palaeobiogeographySauropterygians |journal=JournalPalaeontologia of PalaeogeographyElectronica |volume=822 |issue=1 |pages=Article number 2722.1.1 |doi=10.118626879/s42501913 |s2cid=135461586 |doi-019-0043-5access=free }}</ref>
* Four teeth of a freshwater plesiosaur are described from the [[Middle Jurassic]] [[Xinhe Formation]] ([[Gansu]], [[China]]) by Gao ''et al.'' (2019).<ref>{{cite journal |author1=Ting Gao |author2=Da-Qing Li |author3=Long-Feng Li |author4=Jing-Tao Yang |year=2019 |title=The first record of freshwater plesiosaurian from the Middle Jurassic of Gansu, NW China, with its implications to the local palaeobiogeography |journal=Journal of Palaeogeography |volume=8 |issue=1 |article-number=27 |doi=10.1186/s42501-019-0043-5 |bibcode=2019JPalG...8...27G |s2cid=199547716 |doi-access=free }}</ref>
* The first plesiosaur remains from [[Peru]] are described from the Lower Cretaceous [[La Herradura Formation]] by Meza-Vélez & O'Gorman (2019).<ref>{{cite journal |author1=Iván Meza-Vélez |author2=José P. O'Gorman |year=2019 |title=First plesiosaurian record (Diapsida; Sauropterygia) from the La Herradura Formation, (Valanginian–Hauterivian), Morro Solar, Peru |journal=Cretaceous Research |volume=106 |article-number=104247 |doi=10.1016/j.cretres.2019.104247 |s2cid=204267246 }}</ref>
* A review of the fossil record of Late Cretaceous Antarctic plesiosaurs is published by O'Gorman ''et al.'' (2019).<ref>{{cite journal |author1=José Patricio O'Gorman |author2=Rodrigo Otero |author3=Marcelo Reguero |author4=Zulma Gasparini |year=2019 |title=Cretaceous Antarctic plesiosaurs: stratigraphy, systematics and paleobiogeography |journal=Advances in Polar Science |volume=30 |issue=3 |pages=210–227 |doi=10.13679/j.advps.2018.0049 }}</ref>
* AFossils studyof ona thelarge-bodied morphologypliosaurid-like ofplesiosaur theare teethdescribed andfrom skullthe of[[Middle ''Jurassic]] ([[MegacephalosaurusBajocian]]) eulerti'',[[Passwang andFormation]] on their implications for assessing the phylogenetic relationships of this species, is published([[Switzerland]]) by MadziaSachs, SachsKlug & LindgrenKear (2019).<ref>{{cite journal |author1=DanielSven MadziaSachs |author2=SvenChristian SachsKlug |author3=JohanBenjamin LindgrenP. Kear |year=2019 |title=MorphologicalRare and phylogenetic aspectsevidence of the dentition ofa ''Megacephalosaurusgiant eulerti'',pliosaurid-like a pliosauridplesiosaur from the TuronianMiddle ofJurassic Kansas,(lower USA, with remarks on the cranial anatomyBajocian) of the taxonSwitzerland |journal=Geological[[Swiss MagazineJournal of Palaeontology]] |volume=156138 |issue=72 |pages=1201–1216337–342 |doi=10.10171007/S0016756818000523s13358-019-00200-9 |s2cid=207827907 |doi-access=free }}</ref>
* NewA plesiosaurstudy fossilson arethe describedmorphology fromof the teeth and skull of ''[[BarremianMegacephalosaurus]] levelseulerti'', ofand theon [[Arcillastheir deimplications Morellafor Formation]]assessing ([[Spain]])the byphylogenetic Quesadarelationships ''etof al.''this (2019)species, includingis thepublished firstby [[Leptocleididae|leptocleidid]]Madzia, fossilSachs reported& fromLindgren the [[Iberian Peninsula]](2019).<ref>{{cite journal |author1=J.M.Daniel QuesadaMadzia |author2=A.Sven Pérez-GarcíaSachs |author3=J.M.Johan Gasulla |author4=F. OrtegaLindgren |year=2019 |title=PlesiosauriaMorphological remainsand fromphylogenetic aspects of the Barremiandentition of Morella''Megacephalosaurus (Castellóneulerti'', Spain)a andpliosaurid firstfrom identificationthe Turonian of LeptocleididaeKansas, USA, with inremarks on the Iberiancranial recordanatomy of the taxon |journal=CretaceousGeological ResearchMagazine |volume=94156 |issue=7 |pages=8–241201–1216 |doi=10.10161017/jS0016756818000523 |bibcode=2019GeoM.cretres.2018156.10.0101201M |s2cid=133859507 }}</ref>
* AAn study on the skullisolated [[Morphology (biology)Pliosauridae|morphologypliosaurid]] oftooth twocrown specimensis of ''[[Dolichorhynchops]] bonneri''described from the [[Pierre ShaleHauterivian]] of [[South DakotaAustria]], asby wellLukeneder as& onZverkov (2019), representing the phylogeneticfirst relationshipspliosaur offrom this species,country isand publishedthe bysecond Morganoccurrence &of O'Keefeconical-toothed (2019)pliosaurid in the Hauterivian worldwide.<ref>{{cite journal |author1=DonaldAlexander J. Morgan IIILukeneder |author2=F. RobinNikolay O'KeefeZverkov |year=2019 |title=TheFirst cranial osteologyevidence of twoa specimensconical-toothed of ''Dolichorhynchops bonneri''pliosaurid (PlesiosauriaReptilia, PolycotylidaeSauropterygia) fromin the CampanianHauterivian of Souththe Dakota,Northern andCalcareous a cladistic analysis of theAlps, PolycotylidaeAustria |journal=Cretaceous Research |volume=96106 |pagesarticle-number=149–171104248 |doi=10.1016/j.cretres.20182019.11.027104248 |s2cid=204251536 }}</ref>
* A study on bone [[histology]] and ontogeny of the gravidnew specimen of ''[[PolycotylusStenorhynchosaurus]] latipinnusmunozi'' displayedis atdescribed from the upper [[Los Angeles County Museum of Natural HistoryBarremian]], andof onthe itsArcillolitas implicationsAbigarradas forMember interpretingof athe histological[[Paja growthFormation]] series([[Colombia]]) inby ''Dolichorhynchops[[María bonneri''Páramo|Páramo-Fonseca]], isBenavides-Cabra published& by O’Keefe ''et al.''Gutiérrez (2019).<ref>{{citeCite journal |author1=F.María RE. O’KeefePáramo-Fonseca |author2=P.Cristian MD. SanderBenavides-Cabra |author3=T.Ingry Wintrich |author4=SE. WerningGutiérrez |year=2019 |title=OntogenyA new specimen of polycotylid''Stenorhynchosaurus longmunozi'' bonePáramo-Fonseca microanatomyet al., 2016 (Plesiosauria, Pliosauridae), from the Barremian of Colombia: new morphological features and histologyontogenetic implications |journal=IntegrativeJournal Organismalof BiologyVertebrate Paleontology |volume=139 |issue=14 |pages=oby007e1663426 |doi=10.10931080/iob/oby00702724634.2019.1663426 |s2cid=208561823 }}</ref>
* Description of fossils of plesiosaurs from the Late Jurassic of European [[Russia]] belonging or related to the genus ''[[Colymbosaurus]]'', and a study evaluating the [[Biogeography#Paleobiogeography|palaeobiogeographic]] implications of these fossils, is published online by Arkhangelsky ''et al.'' (2019).<ref>{{cite journal |author1=Maxim S. Arkhangelsky |author2=Nikolay G. Zverkov |author3=Mikhail A. Rogov |author4=Ilya M. Stenshin |author5=Evgeniya M. Baykina |year=2019 |title=Colymbosaurines from the Upper Jurassic of European Russia and their implication for palaeobiogeography of marine reptiles |journal=Palaeobiodiversity and Palaeoenvironments |volume=100 |issue=1 |pages=197–218 |doi=10.1007/s12549-019-00397-0 |s2cid=201983459 }}</ref>
* Skull and neck bones of an [[Elasmosauridae|elasmosaurid]] plesiosaur are described from the [[Cenomanian]] [[Hegushi Formation]] ([[Japan]]) by Utsunomiya (2019), representing the oldest confirmed elasmosaurid in Japan and in East Asia.<ref>{{cite journal |author=Satoshi Utsunomiya |year=2019 |title=Oldest Elasmosauridae(Plesiosauria) in East Asia from the Upper Cretaceous Goshoura Group, Shishijima Island, southwestern Japan |journal=Bulletin of the Osaka Museum of Natural History |volume=73 |pages=23–35 |doi=10.20643/00001333 }}</ref>
* A postcranialstudy skeletonon ofthe anmobility elasmosauridof belongingthe orneck related to the genusof ''[[AristonectesCryptoclidus]] eurymerus'' is described from the uppermost [[Maastrichtian]] levels of the [[Lopez de Bertodano Formation]] ([[Seymour Island]], [[Antarctica]])published by O'GormanWintrich ''et al.'' (2019), who report that this specimen is one of the largest known elasmosaurid specimens worldwide.<ref>{{Citecite journal |author1=J.P.Tanja O'GormanWintrich |author2=S.René SantillanaJonas |author3=R.Hans-Joachim OteroWilke |author4=MLars Schmitz |author5=P. RegueroMartin Sander |year=2019 |title=ANeck giantmobility elasmosauridin (Sauropterygia;the Plesiosauria)Jurassic fromplesiosaur Antarctica''Cryptoclidus eurymerus'': finite element analysis as a new informationapproach onto elasmosauridunderstanding bodythe sizecervical diversityskeleton andin aristonectinefossil evolutionary scenariosvertebrates |journal=Cretaceous ResearchPeerJ |volume=1027 |pagesarticle-number=37–58e7658 |doi=10.10167717/jpeerj.cretres.2019.05.0047658 |pmid=31720095 |pmc=6842296 |doi-access=free }}</ref>
* DigitalNew [[endocast]]splesiosaur offossils twoare specimensdescribed offrom the ''[[LibonectesBarremian]] morgani''levels andof athe [[Polycotylidae|polycotylid]]Arcillas fromde theMorella [[TuronianFormation]] of Goulmima ([[MoroccoSpain]]) are reconstructed by AllemandQuesada ''et al.'' (2019), including the first [[Leptocleididae|leptocleidid]] fossil reported from the [[Iberian Peninsula]].<ref>{{Citecite journal |author1=RémiJ.M. AllemandQuesada |author2=AlexandraA. HoussayePérez-García |author3=NathalieJ.M. BardetGasulla |author4=PeggyF. VincentOrtega |year=2019 |title=EndocranialPlesiosauria remains from the anatomyBarremian of plesiosauriansMorella (ReptiliaCastellón, PlesiosauriaSpain) fromand thefirst Lateidentification Cretaceousof (Turonian)Leptocleididae ofin Goulmimathe (SouthernIberian Morocco)record |journal=Journal of VertebrateCretaceous PaleontologyResearch |volume=39 |issue=294 |pages=e15956368–24 |doi=10.10801016/02724634j.2019cretres.15956362018.10.010 |s2cid=134139253 }}</ref>
* A study on the skull [[Morphology (biology)|morphology]] of two specimens of ''[[Dolichorhynchops]] bonneri'' from the [[Pierre Shale]] of [[South Dakota]], as well as on the phylogenetic relationships of this species, is published by Morgan & O'Keefe (2019).<ref>{{cite journal |author1=Donald J. Morgan III |author2=F. Robin O'Keefe |year=2019 |title=The cranial osteology of two specimens of ''Dolichorhynchops bonneri'' (Plesiosauria, Polycotylidae) from the Campanian of South Dakota, and a cladistic analysis of the Polycotylidae |journal=Cretaceous Research |volume=96 |pages=149–171 |doi=10.1016/j.cretres.2018.11.027 |s2cid=134887820 }}</ref>
* A study on bone [[histology]] and ontogeny of the gravid specimen of ''[[Polycotylus]] latipinnus'' displayed at the [[Los Angeles County Museum of Natural History]], and on its implications for interpreting a histological growth series in ''Dolichorhynchops bonneri'', is published by O'Keefe ''et al.'' (2019).<ref>{{cite journal |author1=F. R. O'Keefe |author2=P. M. Sander |author3=T. Wintrich |author4=S. Werning |year=2019 |title=Ontogeny of polycotylid long bone microanatomy and histology |journal=Integrative Organismal Biology |volume=1 |issue=1 |pages=oby007 |doi=10.1093/iob/oby007 |pmid=33791514 |pmc=7671113 }}</ref>
* Skull and neck bones of an [[Elasmosauridae|elasmosaurid]] plesiosaur are described from the [[Cenomanian]] [[Hegushi Formation]] ([[Japan]]) by Utsunomiya (2019), representing the oldest confirmed elasmosaurid in Japan and in East Asia.<ref>{{cite journal |author=Satoshi Utsunomiya |year=2019 |title=Oldest Elasmosauridae(Plesiosauria) in East Asia from the Upper Cretaceous Goshoura Group, Shishijima Island, southwestern Japan |journal=Bulletin of the Osaka Museum of Natural History |volume=73 |pages=23–35 |doi=10.20643/00001333 }}</ref>
* A postcranial skeleton of an elasmosaurid belonging or related to the genus ''[[Aristonectes]]'' is described from the uppermost [[Maastrichtian]] levels of the [[Lopez de Bertodano Formation]] ([[Seymour Island]], [[Antarctica]]) by O'Gorman ''et al.'' (2019), who report that this specimen is one of the largest known elasmosaurid specimens worldwide.<ref>{{Cite journal|author1=J.P. O'Gorman |author2=S. Santillana |author3=R. Otero |author4=M. Reguero |year=2019 |title=A giant elasmosaurid (Sauropterygia; Plesiosauria) from Antarctica: new information on elasmosaurid body size diversity and aristonectine evolutionary scenarios |journal=Cretaceous Research |volume=102 |pages=37–58 |doi=10.1016/j.cretres.2019.05.004 |s2cid=181725020 }}</ref>
* Description of new fossil material of ''[[Kawanectes]] lafquenianum'' from the upper [[Campanian]]–[[Maastrichtian]] levels of the [[La Colonia Formation]] ([[Argentina]]), extending known stratigraphical range of this taxon and providing new information on diagnostic character states for ''K. lafquenianum'', is published online by O'Gorman (2019).<ref>{{Cite journal|author=José Patricio O'Gorman |year=2019 |title=First record of ''Kawanectes lafquenianum'' (Plesiosauria, Elasmosauridae) from the La Colonia Formation of Argentina, with comments on the mandibular morphology of elasmosaurids |journal=Alcheringa: An Australasian Journal of Palaeontology |volume=44 |issue=1 |pages=176–193 |doi=10.1080/03115518.2019.1687754 |s2cid=213090343 }}</ref>
* Digital [[endocast]]s of two specimens of ''[[Libonectes]] morgani'' and a [[Polycotylidae|polycotylid]] from the [[Turonian]] of Goulmima ([[Morocco]]) are reconstructed by Allemand ''et al.'' (2019).<ref>{{Cite journal|author1=Rémi Allemand |author2=Alexandra Houssaye |author3=Nathalie Bardet |author4=Peggy Vincent |year=2019 |title=Endocranial anatomy of plesiosaurians (Reptilia, Plesiosauria) from the Late Cretaceous (Turonian) of Goulmima (Southern Morocco) |journal=Journal of Vertebrate Paleontology |volume=39 |issue=2 |pages=e1595636 |doi=10.1080/02724634.2019.1595636 |s2cid=181790183 |url=https://hal.archives-ouvertes.fr/hal-02354078/file/Allemand%20et%20al%202019%20Endocranial%20anatomy%20of%20plesiosaurians.pdf }}</ref>
 
===New taxa===
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''[[Cyamodus|Cyamodus orientalis]]'' <ref>{{cite journal |author1=Wei Wang |author2=Chun Li |author3=Torsten M. Scheyer |author4=Lijun Zhao |year=2019 |title=A new species of ''Cyamodus'' (Placodontia, Sauropterygia) from the early Late Triassic of south-west China |journal=Journal of Systematic Palaeontology |volume=17 |issue=17 |pages=1237–12561457–1476 |doi=10.1080/14772019.2018.1535455 |s2cid=91579582 }}</ref>
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Sp. nov
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''[[Glyphoderma|Glyphoderma robusta]]'' <ref>{{cite journal |author1=Jinyuan Hu |author2=Tao Jiang |author3=Zhiguang Li |year=2019 |title=A new species of ''Glyphoderma'' (Reptilia: Placodontia) of Middle Triassic from Fuyuan County, Yunnan Province, China |journal=Journal of Geology |volume=43 |issue=4 |pages=595–598 |url=http://eng.oversea.cnki.net/kcms/detail/detail.aspx?dbCode=cjfq&QueryID=11&CurRec=8&filename=JSDZ201904008&dbname=CJFDLAST2020 |access-date=2020-05-03 |archive-date=2020-09-13 |archive-url=https://web.archive.org/web/20200913000918/http://eng.oversea.cnki.net/kcms/detail/detail.aspx?dbCode=cjfq&QueryID=11&CurRec=8&filename=JSDZ201904008&dbname=CJFDLAST2020 |url-status=dead }}</ref>
''[[Leivanectes]]''<ref>{{cite journal |author1=Maria Eurídice Páramo-Fonseca |author2=José Patricio O'Gorman |author3=Zulma Gasparini |author4=Santiago Padilla |author5=Mary Luz Parra Ruge |year=2019 |title=A new late Aptian elasmosaurid from the Paja Formation, Villa de Leiva, Colombia |journal=Cretaceous Research |volume=99 |pages=30–40 |doi=10.1016/j.cretres.2019.02.010 }}</ref>
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Sp. nov
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Valid
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Hu, Jiang & Li
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[[Middle Triassic]] ([[Ladinian]])
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[[Falang Formation]]
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{{Flag|China}}
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''[[Leivanectes]]''<ref>{{cite journal |author1=Maria Eurídice Páramo-Fonseca |author2=José Patricio O'Gorman |author3=Zulma Gasparini |author4=Santiago Padilla |author5=Mary Luz Parra Ruge |year=2019 |title=A new late Aptian elasmosaurid from the Paja Formation, Villa de Leiva, Colombia |journal=Cretaceous Research |volume=99 |pages=30–40 |doi=10.1016/j.cretres.2019.02.010 |hdl=11336/127809 |s2cid=134171636 |hdl-access=free }}</ref>
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Gen. et sp. nov
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''[[Lindwurmia]]''<ref>{{cite journal |author1=Peggy Vincent |author2=Glenn W. Storrs |year=2019 |title=''Lindwurmia'', a new genus of Plesiosauria (Reptilia: Sauropterygia) from the earliest Jurassic of Halberstadt, northwest Germany |journal=The Science of Nature |volume=106 |issue=1–2 |pagesarticle-number=Article 5 |doi=10.1007/s00114-018-1600-y |pmid=30689058 |bibcode=2019SciNa.106....5V |s2cid=59304744 }}</ref>
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Gen. et sp. nov
Line 377 ⟶ 585:
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An early member of [[Plesiosauria]]. Genus includes new species ''L. thiuda''.
|[[File:Museum_Heineanum_Rudersaurier_Eurycleidus_arcuatus.jpg|center|frameless]]
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''[[Microcleidus|Microcleidus melusinae]]'' <ref>{{cite journal |author1=Peggy Vincent |author2=Robert Weis |author3=Guy Kronz |author4=Dominique Delsate |year=2019 |title=''Microcleidus melusinae'', a new plesiosaurian (Reptilia, Plesiosauria) from the Toarcian of Luxembourg |journal=Geological Magazine |volume=156 |issue=1 |pages=99–116 |doi=10.1017/S0016756817000814 |bibcode=2019GeoM..156...99V |s2cid=135111068 |url=https://hal.sorbonne-universite.fr/hal-02298448/file/Vincent%20et%20al.%20-%202019%20-%20Microcleidus%20melusinae%2C%20a%20new%20plesiosaurian%20%28Repti.pdf }}</ref>
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Sp. nov
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''[[Nothosaurus|Nothosaurus cristatus]]''<ref>{{Cite journal|author1=Juliane K. Hinz |author2=Andreas T. Matzke |author3=Hans-Ulrich Pfretzschner |year=2019 |title=A new nothosaur (Sauropterygia) from the Ladinian of Vellberg-Eschenau, southern Germany |journal=Journal of Vertebrate Paleontology |volume=39 |issue=2 |pages=e1585364 |doi=10.1080/02724634.2019.1585364 |s2cid=155775644 }}</ref>
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Sp. nov
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''[[Panzhousaurus]]''<ref>{{cite journal |author1=Da-Yong Jiang |author2=Wen-Bin Lin |author3=Olivier Rieppel |author4=Ryosuke Motani |author5=Zuo-Yu Sun |year=2019 |title=A new Anisian (Middle Triassic) eosauropterygian (Reptilia, Sauropterygia) from Panzhou, Guizhou Province, China |journal=Journal of Vertebrate Paleontology |volume=38 |issue=4 |pages=(1)–(9) |doi=10.1080/02724634.2018.1480113 |s2cid=109794338 }}</ref>
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Gen. et sp. nov
Line 434 ⟶ 642:
==Turtles==
===Research===
* A study on the phylogenetic relationships of living and fossil turtles is published by Evers & Benson (2019).<ref>{{cite journal |author1=Serjoscha W. Evers |author2=Roger B. J. Benson |year=2019 |title=A new phylogenetic hypothesis of turtles with implications for the timing and number of evolutionary transitions to marine lifestyles in the group |journal=Palaeontology |volume=62 |issue=1 |pages=93–134 |doi=10.1111/pala.12384 |s2cid=134736808 |url=https://ora.ox.ac.uk/objects/uuid:33a26ae9-a5f6-43b5-ace1-2d17b1dadeb6 }}</ref>
* A study on the evolution and [[Ontogeny|ontogenetic]] development of the akinetic skull of turtles, based on data from extant and fossil taxa, is published by Werneburg & Maier (2019).<ref>{{cite journal |author1=Ingmar Werneburg |author2=Wolfgang Maier |year=2019 |title=Diverging development of akinetic skulls in cryptodire and pleurodire turtles: an ontogenetic and phylogenetic study |journal=Vertebrate Zoology |volume=69 |issue=2 |pages=113–143 |doi=10.26049/VZ69-2-2019-01 }}</ref>
* A study on the shell composition in [[Proterochersidae|proterochersidshistology]] of shell bones of extant and otherfossil [[Triassic]]turtles, [[Pantestudines|pantestudinates]]evaluating its utility for determination of habitat of fossil turtles, is published online by SzczygielskiJannello, Cerda & Sulejde la Fuente (2019).<ref>{{citeCite journal |author1=TomaszJuan M. SzczygielskiJannello |author2=TomaszIgnacio SulejA. Cerda |author3=Marcelo S. de la Fuente |year=2019 |title=The earlyrelationship compositionbetween andbone evolutionshell ofmicroanatomy theand turtlepalaeoecology shell (Reptilia,in Testudinata) from South America |journal=PalaeontologyPalaeogeography, Palaeoclimatology, Palaeoecology |volume=62537 |issuearticle-number=3 |pages=375–415109412 |doi=10.11111016/palaj.12403palaeo.2019.109412 |s2cid=210293162 }}</ref>
* A study on the shell composition in [[Proterochersidae|proterochersids]] and other [[Triassic]] [[Pantestudines|pantestudinates]] is published by Szczygielski & Sulej (2019).<ref>{{cite journal |author1=Tomasz Szczygielski |author2=Tomasz Sulej |year=2019 |title=The early composition and evolution of the turtle shell (Reptilia, Testudinata) |journal=Palaeontology |volume=62 |issue=3 |pages=375–415 |doi=10.1111/pala.12403 |s2cid=134011993 }}</ref>
* Fragmentary fossil material of a [[Basal (phylogenetics)|basal]] turtle belonging to the clade [[Mesochelydia]] will beis described from the [[Lower Cretaceous]] Teete locality ([[Sakha Republic|Yakutia]], [[Russia]]) by Skutschas ''et al.'' (2019).<ref>{{cite journal |author1=Pavel P. Skutschas |author2=Valentina D. Markova |author3=Veniamin V. Kolchanov |author4=Alexander O. Averianov |author5=Thomas Martin |author6=Rico Schellhorn |author7=Petr N. Kolosov |author8=Dmitry V. Grigoriev |author9=Dmitry D. Vitenko |author10=Ekaterina M. Obraztsova |author11=Igor G. Danilov |year=2019 |title=Basal turtle material from the Lower Cretaceous of Yakutia (Russia) filling the gap in the Asian record |journal=Cretaceous Research |volume=in press106 |pagesarticle-number=104186 |doi=10.1016/j.cretres.2019.07.016 |s2cid=202195840 }}</ref>
* Description of new fossil material of ''[[Condorchelys]] antiqua'', and a study on the phylogenetic relationships of early turtles, is published by Sterli, de la Fuente & Rougier (2019).<ref>{{cite journal |author1=Juliana Sterli |author2=Marcelo S. de la Fuente |author3=Guillermo W. Rougier |year=2019 |title=New remains of ''Condorchelys antiqua'' (Testudinata) from the Early-Middle Jurassic of Patagonia: anatomy, phylogeny, and paedomorphosis in the early evolution of turtles |journal=Journal of Vertebrate Paleontology |volume=38 |issue=4 |pages=(1)–(17) |doi=10.1080/02724634.2018.1480112 |s2cid=109556104 |hdl=11336/99525 |hdl-access=free }}</ref>
* DescriptionA ofstudy newon the fossilendocranial materialanatomy of ''[[PeligrochelysNaomichelys]] walshaespeciosa'' from the [[Paleocene]] ([[Danian]]) [[Salamanca Formation]] ([[Argentina]]), and a study on the phylogenetic relationships of this species, is published by Paulina-Carabajal, Sterli & de la FuenteWerneburg (2019).<ref>{{cite journal |author1=JulianaAriana SterliPaulina-Carabajal |author2=MarceloJuliana S.Sterli de la|author3=Ingmar FuenteWerneburg |year=2019 |title=CranialThe andendocranial post-cranial remains and phylogenetic relationshipsanatomy of the Gondwanan meiolaniformstem turtle ''PeligrochelysNaomichelys walshaespeciosa'' from the PaleoceneEarly Cretaceous of Chubut,North ArgentinaAmerica |journal=JournalActa ofPalaeontologica PaleontologyPolonica |volume=9364 |issue=4 |pages=798–821711–716 |doi=10.10174202/jpaapp.201900606.112019 |doi-access=free |hdl=11336/175025 |hdl-access=free }}</ref>
* TheDescription onlyof completenew shellfossil material of ''[[EocenochelusPeligrochelys]] walshae'' identified so far will be described from the [[EocenePaleocene]] ([[LutetianDanian]]) [[SobrarbeSalamanca Formation]] ([[SpainArgentina]]), and a study on the phylogenetic relationships of this species, is published by Pérez-GarcíaSterli ''et& al.''de la Fuente (2019).<ref>{{cite journal |author1=A.Juliana Pérez-GarcíaSterli |author2=E.Marcelo Díaz-Berenguer |author3=AS. Badiolade |author4=J.I.la CanudoFuente |year=2019 |title=AnCranial unexpectedand finding:post-cranial identificationremains ofand thephylogenetic first complete shellrelationships of the Franco-Belgian middle Eocene littoralGondwanan pleurodiranmeiolaniform turtle ''EocenochelusPeligrochelys erembertiwalshae'' infrom Spainthe |journal=HistoricalPaleocene Biology:of AnChubut, InternationalArgentina |journal=Journal of PaleobiologyPaleontology |volume=in press93 |issue=4 |pages=798–821 1–7|doi=10.10801017/08912963jpa.2019.164433011 |s2cid=146608711 }}</ref>
* DescriptionThe ofonly newcomplete fossil materialshell of ''[[PlesiochelysEocenochelus]] bigleri'' identified so far is described from the [[KimmeridgianEocene]] Banné Marls ([[SwitzerlandLutetian]]), providing[[Sobrarbe newFormation]] information on the anatomy of this species, is published([[Spain]]) by RaselliPérez-García &''et Anquetinal.'' (2019).<ref>{{Citecite journal |author1=IrenaA. RaselliPérez-García |author2=JérémyE. Díaz-Berenguer |author3=A. Badiola |author4=J.I. AnquetinCanudo |year=2019 |title=NovelAn insightsunexpected intofinding: identification of the morphologyfirst complete shell of the Franco-Belgian middle Eocene littoral pleurodiran turtle ''PlesiochelysEocenochelus biglerieremberti'' fromin theSpain early|journal=Historical KimmeridgianBiology: ofAn NorthwesternInternational SwitzerlandJournal |journal=PLoSof ONEPaleobiology |volume=1433 |issue=54 |pages=e0214629527–533 |doi=10.13711080/journal08912963.pone2019.0214629 |pmid=310912411644330 |pmcs2cid=6519798199637318 }}</ref>
* RedescriptionDescription of thenew [[holotype]]fossil specimenmaterial of ''[[NanhsiungchelysNeusticemys]] wuchingensisneuquina'' from the Upper Jurassic of the Neuquén Basin ([[Argentina]]) and a study on the phylogenetic relationships of this species is published online by TongRuiz, de la Fuente & LiFernández (2019).<ref>{{cite journal |author1=HaiyanP. TongGonzález Ruiz |author2=LuM. S. de la Fuente |author3=M.S. LiFernández |year=2019 |title=ANew revisioncranial fossils of the holotypeJurassic ofturtle ''NanhsiungchelysNeusticemys wuchingensisneuquina'', Ye,and 1966phylogenetic (Testudines:relationships Cryptodira:of Trionychoidae:the Nanhsiungchelyidae)only thalassochelydian known from the eastern Pacific |journal=CretaceousJournal of ResearchPaleontology |volume=9594 |issue=1 |pages=151–163145–164 |doi=10.10161017/jjpa.cretres.20182019.11.00374 |s2cid=210611801 }}</ref>
* A reviewDescription of thenew fossil material of ''[[Araripemydidae|araripemydidPlesiochelys]] fossil recordbigleri'' from Africathe is[[Kimmeridgian]] publishedBanné by Pérez-GarcíaMarls (2019[[Switzerland]]), whoproviding considersnew ''Laganemysinformation tenerensis''on tothe beanatomy aof [[Synonymthis (taxonomy)|juniorspecies, synonym]]is ofpublished ''[[Taquetochelys]]by decorata''Raselli & Anquetin (2019).<ref>{{citeCite journal|author1=Irena Raselli |authorauthor2=AdánJérémy Pérez-GarcíaAnquetin |year=2019 |title=IdentificationNovel ofinsights into the Lowermorphology Cretaceous pleurodiran turtleof ''TaquetochelysPlesiochelys decoratabigleri'' asfrom the onlyearly AfricanKimmeridgian araripemydidof speciesNorthwestern Switzerland |journal=Comptes RendusPLOS PalevolONE |volume=1814 |issue=15 |pages=24–32e0214629 |doi=10.10161371/jjournal.crpvpone.20180214629 |pmid=31091241 |pmc=6519798 |bibcode=2019PLoSO.04.0041414629R |doi-access=free }}</ref>
* A studyshell onof the''Plesiochelys anatomybigleri'' ofwhich themight shellhave ofbeen thetrodden on by a large [[BothremydidaeSauropoda|bothremydidsauropod]] speciesdinosaur, representing the first evidence that these turtles occasionally visited ''[[Taphrosphystidal flat]] congolensis''environments, and on itsis implicationsreported for inferringfrom the taxonomicLate compositionJurassic of thePorrentruy genus(Swiss ''Taphrosphys'',[[Jura will be publishedMountains]]) by PérezPüntener García,''et Mees & Smithal.'' (2019).<ref>{{citeCite journal |author1=AdánChristian Pérez GarcíaPüntener |author2=FloriasJean-Paul MeesBillon-Bruyat |author3=ThierryDaniel Marty |author4=Géraldine SmithParatte |year=2019 |title=ShellUnder anatomythe feet of thesauropods: Africana Paleocenetrampled bothremydidcoastal marine turtle ''Taphrosphysfrom congolensis''the andLate systematicJurassic implicationsof within TaphrosphyiniSwitzerland? |journal=Historical Biology: An International[[Swiss Journal of PaleobiologyGeosciences]] |volume=in press112 |issue=2–3 |pages=507–515 1–10|doi=10.10801007/08912963s00015-019-00347-0 |s2cid=202565198 |url=http://osf.2018.1497023io/2atnq/ }}</ref>
* DescriptionRedescription of fossils of ''[[Ordosemys]] leios'' from the Lower Cretaceous [[Mengyin Formationholotype]] ([[China]]), and a study on their implications for inferring the ecologyspecimen of this species and the age of the ''[[Luohandong FormationNanhsiungchelys]] of the [[Ordos Basin]],wuchingensis'' is published by LiTong ''et& al.''Li (2019).<ref>{{cite journal |author1=Da-QingHaiyan LiTong |author2=Chang-Fu Zhou |author3=LanLu Li |author4=Jing-Tao Yang |author5=Longfeng Li |author6=Márton Rabi |year=2019 |title=TheA sinemydidrevision turtle ''Ordosemys'' fromof the Lowerholotype Cretaceousof Mengyin''Nanhsiungchelys Formationwuchingensis'', of ShandongYe, China1966 and(Testudines: itsCryptodira: implicationTrionychoidae: for the age of the Luohandong Formation of the Ordos BasinNanhsiungchelyidae) |journal=PeerJCretaceous Research |volume=795 |pages=e6229151–163 |doi=10.77171016/peerjj.6229 |pmid=30671300cretres.2018.11.003 |pmcs2cid=6338100133937906 }}</ref>
* A study on shifts of rangereview of the [[European pond turtleAraripemydidae|araripemydid]] infossil Easternrecord Europefrom throughoutAfrica theis [[Holocene]],published basedby onPérez-García data(2019), fromwho [[subfossil]]considers remains''Laganemys fromtenerensis'' archaeologicalto sites,be isa published[[Synonym by(taxonomy)|junior Nekrasovasynonym]] of ''et[[Taquetochelys]] al.decorata'' (2019).<ref>{{Citecite journal|author1=Oksana Nekrasova |author2author=YevheniiaAdán Yanish |author3=Volodymyr Tytar |author4=Mihails PupinsPérez-García |year=2019 |title=GIS-modelingIdentification of the rangeLower shiftsCretaceous of the sub-fossil and extant European pondpleurodiran turtle (''EmysTaquetochelys orbicularisdecorata'') inas the Easternonly EuropeAfrican inararipemydid Holocenespecies |journal=DiversityComptes Rendus Palevol |volume=1118 |issue=81 |pages=Article 12124–32 |doi=10.33901016/d11080121j.crpv.2018.04.004 |bibcode=2019CRPal..18...24P |s2cid=134080281 }}</ref>
* A laterevision [[Pleistocene]]of nuchalthe bonefossil isrecord described fromof the Muaco site (western non-[[VenezuelaBaenidae|baenid]]) by Cadena & Carrillo-Briceño (2019), who interpret this specimen as the first undisputable fossil of a membermembers of the genusclade ''[[RhinoclemmysParacryptodira]]'' foundis eastpublished ofby theJoyce [[Andes]]& Anquetin (2019).<ref>{{cite journal |author1=EdwinWalter AlbertoG. CadenaJoyce |author2=JorgeJérémy D. Carrillo-BriceñoAnquetin |year=2019 |title=FirstA fossilreview of ''Rhinoclemmys''the Fitzinger,fossil 1826record (Cryptodira,of Geoemydidae)nonbaenid eastturtles of the Andesclade Paracryptodira |journal=SouthBulletin of the AmericanPeabody JournalMuseum of HerpetologyNatural History |volume=1460 |issue=12 |pages=19–23129–155 |doi=10.29943374/SAJH-D-17-00099014.1060.0204 |s2cid=203780510 |url=http://osf.io/8v6mx/ }}</ref>
* ADescription studyof on the massfossils of North American Pleistocene ''[[tortoiseOrdosemys]]s, onleios'' from the relationshipLower betweenCretaceous the[[Mengyin massFormation]] of tortoises([[China]]), and their ability to maintain a viablestudy bodyon temperaturetheir atimplications lowfor ambient temperatures, and oninferring the implicationsecology of this relationshipsspecies forand the knowledgeage of Pleistocene temperatures in the areas[[Luohandong fromFormation]] whichof tortoisethe fossils[[Ordos are knownBasin]], is published by Esker,Li Forman''et & Butleral.'' (2019).<ref>{{cite journal |author1=DonaldDa-Qing A. EskerLi |author2=SteveChang-Fu L. FormanZhou |author3=DavaLan K.Li |author4=Jing-Tao Yang |author5=Longfeng Li |author6=Márton ButlerRabi |year=2019 |title=ReconstructingThe sinemydid turtle ''Ordosemys'' from the massLower Cretaceous Mengyin Formation of Shandong, China and thermalits ecologyimplication for the age of Norththe AmericanLuohandong Formation of the PleistoceneOrdos tortoisesBasin |journal=PaleobiologyPeerJ |volume=457 |issuearticle-number=2 |pages=363–377e6229 |doi=10.10177717/pabpeerj.2019.66229 |pmid=30671300 |pmc=6338100 |doi-access=free }}</ref>
* AFossil studyspecimens onbelonging to the skeletalspecies anatomy''Ordosemys andliaoxiensis'', phylogeneticotherwise relationshipsknown from the older [[Yixian Formation]] of ''the [[RhinochelysJehol Biota]], pulchriceps''are isdescribed publishedfrom the Lower Cretaceous [[Hengtongshan Formation]] ([[Jilin]], [[China]]) by EversZhou, BarrettWu & BensonRabi (2019).<ref>{{citeCite journal |author1=SerjoschaChang-Fu W. EversZhou |author2=PaulWen-Hao M. BarrettWu |author3=RogerMárton B. J. BensonRabi |year=2019 |title=AnatomyPresence of the Jehol Biota turtle ''RhinochelysOrdosemys pulchricepsliaoxiensis'' (Protostegidae)in andthe marineEarly adaptationCretaceous duringHengtongshan theFormation earlyof evolutionsouthern ofJilin chelonioidsProvince, China |journal=PeerJFossil Record |volume=722 |issue=2 |pages=e681157–64 |doi=10.77175194/peerj.6811fr-22-57-2019 |pmids2cid=31106054203380800 |pmcdoi-access=6500378free }}</ref>
* DigitalA endocastsstudy ofon theshifts brainof cavity and endosseous labyrinthrange of ''Rhinochelysthe pulchriceps''[[European willpond beturtle]] presentedin byEastern EversEurope ''etthroughout al.''the (2019)[[Holocene]], whobased useon thesedata endocasts to studyfrom [[neuroanatomysubfossil]] andremains [[carotid]]from circulationarchaeological ofsites, thisis speciespublished by Nekrasova ''et al.'' (2019).<ref>{{citeCite journal |author1=Serjoscha W.Oksana EversNekrasova |author2=JamesYevheniia M. NeenanYanish |author3=GabrielVolodymyr S. FerreiraTytar |author4=Ingmar Werneburg |author5=Paul M. Barrett |author6=Roger B. J.Mihails BensonPupins |year=2019 |title=Neurovascular anatomyGIS-modeling of the protostegidrange turtleshifts ''Rhinochelysof pulchriceps''the sub-fossil and comparisonsextant ofEuropean membranouspond andturtle endosseous(''Emys labyrinth shapeorbicularis'') in anEastern extantEurope turtlein Holocene |journal=Zoological Journal of the Linnean SocietyDiversity |volume=in press11 |issue=8 |pagesarticle-number=121 |doi=10.10933390/zoolinnean/zlz063d11080121 |doi-access=free }}</ref>
* A gravidlate specimen of ''[[Desmatochelys padillaiPleistocene]]'', representingnuchal the first indisputable gravid marine fossil turtle reported so far,bone is described from the [[LowerMuaco Cretaceous]]site of(western [[ColombiaVenezuela]]) by Cadena ''et& al.''Carrillo-Briceño (2019), who interpret this specimen as indicatingthe thatfirst ''D. padillai'' produced rigid eggs similar to those associated with some extant andundisputable fossil freshwaterof anda terrestrialmember turtles,of andthe unlikegenus flexible''[[Rhinoclemmys]]'' eggsfound producedeast byof extantthe marine turtles[[Andes]].<ref>{{Citecite journal |author1=Edwin‐AlbertoEdwin Alberto Cadena |author2=Mary L. Parra‐Ruge |author3=Juan deJorge D. Parra‐Ruge |author4=Santiago Padilla‐BernalCarrillo-Briceño |year=2019 |title=A gravidFirst fossil turtleof from''Rhinoclemmys'' theFitzinger, Early1826 Cretaceous(Cryptodira, revealsGeoemydidae) aeast differentof eggthe developmentAndes strategy|journal=South toAmerican thatJournal of extant marine turtles |journal=PalaeontologyHerpetology |volume=6214 |issue=41 |pages=533–54519–23 |doi=10.11112994/palaSAJH-D-17-00099.124131 |s2cid=146111011 }}</ref>
* A specimenstudy on the mass of ''Desmatochelys''North belongingAmerican orPleistocene related[[tortoise]]s, toon the speciesrelationship ''D.between lowii''the ismass describedof fromtortoises theand lowertheir [[Campanian]]ability [[Austinto Formation]]maintain ([[Coahuila]],a [[Mexico]])viable bybody López-Condetemperature ''etat al.''low (2019)ambient temperatures, representingand on the first recordimplications of this relationships for the familyknowledge [[Protostegidae]]of Pleistocene temperatures in the Lateareas Cretaceousfrom ofwhich Mexicotortoise reportedfossils soare farknown, is published by Esker, Forman & Butler (2019).<ref>{{Citecite journal |author1=OliverDonald A. López-CondeEsker |author2=JulianaSteve SterliL. Forman |author3=JesúsDava Alvarado-Ortega |author4=María LK. Chavarría-Arellano |author5=Héctor Porras-MúzquizButler |year=2019 |title=TheReconstructing firstthe recordmass ofand ''Desmatochelys''thermal cf.ecology ''D.of lowii''North fromAmerican thePleistocene Late Cretaceous (Campanian) of Coahuila, Mexicotortoises |journal=Journal of South American Earth SciencesPaleobiology |volume=9445 |issue=2 |pages=Article 102204363–377 |doi=10.10161017/j.jsamespab.2019.05.0206 |s2cid=155216574 }}</ref>
* A study on the phylogenetic relationships and biogeographic origin of tortoises belonging to the genus ''[[Cylindraspis]]'', based on data from near-complete mitochondrial genomes, is published by Kehlmaier ''et al.'' (2019).<ref>{{Cite journal|author1=Christian Kehlmaier |author2=Eva Graciá |author3=Patrick D. Campbell |author4=Margaretha D. Hofmeyr |author5=Silke Schweiger |author6=Albert Martínez-Silvestre |author7=Walter Joyce |author8=Uwe Fritz |year=2019 |title=Ancient mitogenomics clarifies radiation of extinct Mascarene giant tortoises (''Cylindraspis'' spp.) |journal=Scientific Reports |volume=9 |issue=1 |article-number=17487 |doi=10.1038/s41598-019-54019-y |pmid=31767921 |pmc=6877638 |bibcode=2019NatSR...917487K }}</ref>
* Three incomplete shells of ''[[Protestudo]] bessarabica'' are described from the late Miocene of the Belka locality by Syromyatnikova ''et al.'' (2019), representing the first record of this species from [[Ukraine]] reported so far.<ref>{{Cite journal|author1=E.V. Syromyatnikova |author2=T.V. Krakhmalnaya |author3=V.M. Chkhikvadze |author4=I.G. Danilov |year=2019 |title=Turtles ''Protestudo bessarabica'' from the Late Miocene of Ukraine |journal=Paleontological Journal |volume=53 |issue=6 |pages=647–659 |doi=10.1134/S003103011906011X |s2cid=210926442 |url=https://elibrary.ru/item.asp?id=41169169 |url-access=subscription }}</ref>
* A study on the skeletal anatomy and phylogenetic relationships of ''[[Rhinochelys]] pulchriceps'' is published by Evers, Barrett & Benson (2019).<ref>{{cite journal |author1=Serjoscha W. Evers |author2=Paul M. Barrett |author3=Roger B. J. Benson |year=2019 |title=Anatomy of ''Rhinochelys pulchriceps'' (Protostegidae) and marine adaptation during the early evolution of chelonioids |journal=PeerJ |volume=7 |article-number=e6811 |doi=10.7717/peerj.6811 |pmid=31106054 |pmc=6500378 |doi-access=free }}</ref>
* Digital endocasts of the brain cavity and endosseous labyrinth of ''Rhinochelys pulchriceps'' are presented by Evers ''et al.'' (2019), who use these endocasts to study [[neuroanatomy]] and [[carotid]] circulation of this species.<ref>{{cite journal |author1=Serjoscha W. Evers |author2=James M. Neenan |author3=Gabriel S. Ferreira |author4=Ingmar Werneburg |author5=Paul M. Barrett |author6=Roger B. J. Benson |year=2019 |title=Neurovascular anatomy of the protostegid turtle ''Rhinochelys pulchriceps'' and comparisons of membranous and endosseous labyrinth shape in an extant turtle |journal=Zoological Journal of the Linnean Society |volume=187 |issue=3 |pages=800–828 |doi=10.1093/zoolinnean/zlz063 |doi-access=free }}</ref>
* A gravid specimen of ''[[Desmatochelys padillai]]'', representing the first indisputable gravid marine fossil turtle reported so far, is described from the [[Lower Cretaceous]] of [[Colombia]] by Cadena ''et al.'' (2019), who interpret this specimen as indicating that ''D. padillai'' produced rigid eggs similar to those associated with some extant and fossil freshwater and terrestrial turtles, and unlike flexible eggs produced by extant marine turtles.<ref>{{Cite journal|author1=Edwin-Alberto Cadena |author2=Mary L. Parra-Ruge |author3=Juan de D. Parra-Ruge |author4=Santiago Padilla-Bernal |year=2019 |title=A gravid fossil turtle from the Early Cretaceous reveals a different egg development strategy to that of extant marine turtles |journal=Palaeontology |volume=62 |issue=4 |pages=533–545 |doi=10.1111/pala.12413 |s2cid=134446621 }}</ref>
* A specimen of ''Desmatochelys'' belonging or related to the species ''D. lowii'' is described from the lower [[Campanian]] [[Austin Formation]] ([[Coahuila]], [[Mexico]]) by López-Conde ''et al.'' (2019), representing the first record of the family [[Protostegidae]] in the Late Cretaceous of Mexico reported so far.<ref>{{Cite journal|author1=Oliver A. López-Conde |author2=Juliana Sterli |author3=Jesús Alvarado-Ortega |author4=María L. Chavarría-Arellano |author5=Héctor Porras-Múzquiz |year=2019 |title=The first record of ''Desmatochelys'' cf. ''D. lowii'' from the Late Cretaceous (Campanian) of Coahuila, Mexico |journal=Journal of South American Earth Sciences |volume=94 |article-number=102204 |doi=10.1016/j.jsames.2019.05.020 |bibcode=2019JSAES..9402204L |s2cid=182226555 }}</ref>
* An isolated [[Turtle shell#Carapace|carapacial]] ossicle of a member of the genus ''[[Psephophorus]]'' is described from the lowermost [[Pliocene]] [[Purisima Formation]] ([[California]], [[United States]]) by Fallon & Boessenecker (2019), representing the first occurrence of a sea turtle from this formation.<ref>{{Cite journal|author1=Bailey R. Fallon |author2=Robert W. Boessenecker |year=2019 |title=First record of the leatherback sea turtle (Dermochelyidae) from the Mio-Pliocene Purisima Formation of northern California, USA |journal=PaleoBios |volume=36 |pages=ucmp_paleobios_44240 |url=https://escholarship.org/uc/item/2c38b585 }}</ref>
* DescriptionAn incomplete skeleton of thea anatomyjuvenile ofsea theturtle braincasebelonging ofto athe specimen ofgenus ''[[SyllomusEochelone]] aegyptiacus'' is described from the [[MioceneEocene]] ([[Calvert FormationBartonian]]) of the Gorny Luch locality ([[VirginiaKrasnodar Krai]], [[United StatesRussia]]) is published by MatzkeZvonok &''et Maischal.'' (2019).<ref>{{Cite journal|author1=Andreas TE.A. MatzkeZvonok |author2=MichaelE.V. WSyromyatnikova |author3=I.G. Danilov |author4=A.F. MaischBannikov |year=2019 |title=TheA braincasesea of ''Syllomus aegyptiacus'' Lydekker, 1899turtle (Reptilia, TestudinesCheloniidae) from the Middlemiddle Miocene Calvert FormationEocene of VirginiaNorth Caucasus |journal=PalaeodiversityPaleontological Journal |volume=1253 |issue=15 |pages=31–39530–539 |doi=10.184761134/paleS0031030119050137 |s2cid=203848066 |url=https://elibrary.v12ru/item.a3asp?id=39324160 |url-access=subscription }}</ref>
* Description of turtlethe fossilsanatomy fromof fivethe braincase of a specimen of ''[[PaleogeneSyllomus]] localitiesaegyptiacus'' infrom the [[CrimeaMiocene]] [[Calvert Formation]] ([[Virginia]], [[United States]]) is published by ZvonokMatzke & DanilovMaisch (2019).<ref>{{Cite journal|author1=E.AAndreas T. ZvonokMatzke |author2=I.GMichael W. DanilovMaisch |year=2019 |title=PaleogeneThe turtlesbraincase of Crimea''Syllomus aegyptiacus'' Lydekker, 1899 (Reptilia, Testudines) from the Middle Miocene Calvert Formation of Virginia |journal=Paleontological JournalPalaeodiversity |volume=5312 |issue=1 |pages=62–7331–39 |doi=10.113418476/S003103011901012Xpale.v12.a3 |urls2cid=https://elibrary.ru/item.asp?id155517282 |doi-access=36899662free }}</ref>
* ADescription study onof turtle remainsfossils from five [[HolocenePaleogene]] localities in the [[Thailand|ThaiCrimea]] central plain, and on their implications for the knowledge of changes of turtle biodiversity in this area over the Holocene, is published by ClaudeZvonok ''et& al.''Danilov (2019).<ref>{{Cite journal|author1=JulienE.A. ClaudeZvonok |author2=PrasitI.G. Auetrakulvit |author3=Wilailuck Naksri |author4=Corentin Bochaton |author5=Valéry Zeitoun |author6=HaiyanTongDanilov |year=2019 |title=ThePaleogene recent fossil turtle recordturtles of the central plain of Thailand reveals local extinctionsCrimea |journal=Annales dePaleontological PaléontologieJournal |volume=10553 |issue=1 |pages=62–73 |doi=10.10161134/jS003103011901012X |s2cid=181673636 |url=https://elibrary.annpalru/item.2019.04.005asp?id=36899662 |url-access=subscription }}</ref>
* A study on turtle remains from five [[Holocene]] localities in [[Thailand|Thai]] central plain, and on their implications for the knowledge of changes of turtle biodiversity in this area over the Holocene, is published by Claude ''et al.'' (2019).<ref>{{Cite journal|author1=Julien Claude |author2=Prasit Auetrakulvit |author3=Wilailuck Naksri |author4=Corentin Bochaton |author5=Valéry Zeitoun |author6=HaiyanTong |year=2019 |title=The recent fossil turtle record of the central plain of Thailand reveals local extinctions |journal=Annales de Paléontologie |volume=105 |issue=4 |pages=305–315 |doi=10.1016/j.annpal.2019.04.005 |s2cid=197569669 }}</ref>
 
===New taxa===
Line 472 ⟶ 688:
|-
|
''[[BanhxeochelysAsmodochelys]]''<ref>{{Citecite journal |author1=RafaellaAndrew CD. GarbinGentry |author2=MadelaineJun BöhmeA. Ebersole |author3=WalterCaitlin GR. JoyceKiernan |year=2019 |title=A''Asmodochelys parhami'', a new testudinoidfossil marine turtle from the middleCampanian toDemopolis lateChalk Eoceneand the stratigraphic congruence of Vietnamcompeting marine turtle phylogenies |journal=PeerJRoyal Society Open Science |volume=76 |pagesissue=e628012 |article-number=191950 |doi=10.77171098/peerjrsos.6280191950 |pmid=3080524531903219 |pmc=63835596936288 |bibcode=2019RSOS....691950G }}</ref>
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Gen. et sp. nov
Line 478 ⟶ 694:
Valid
|
Gentry, Ebersole & Kiernan
Garbin, Böhme & Joyce
|
[[Late Cretaceous]] ([[Campanian]])
|
[[Demopolis Chalk]]
|
{{Flag|United States}}<br>({{Flag|Alabama}}<br>{{Flag|Mississippi}})
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A member of the family [[Ctenochelyidae]]. The type species is ''A. parhami''.
|
|-
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''[[Axestemys|Axestemys infernalis]]''<ref>{{cite journal |author1=Walter G. Joyce |author2=Donald B. Brinkman |author3=Tyler R. Lyson |year=2019 |title=A new species of trionychid turtle, ''Axestemys infernalis'' sp. nov., from the Late Cretaceous (Maastrichtian) Hell Creek and Lance formations of the Northern Great Plains, USA |journal=Palaeontologia Electronica |volume=22 |issue=3 |pages=Article number 22.3.72 |doi=10.26879/949 |s2cid=214538408 |doi-access=free }}</ref>
|
Sp. nov
|
Valid
|
Joyce, Brinkman & [[Tyler Lyson|Lyson]]
|
[[Late Cretaceous]] ([[Maastrichtian]])
|
[[Hell Creek Formation]]<br>
[[Lance Formation]]
|
{{Flag|United States}}<br>({{Flag|Montana}}<br>{{Flag|North Dakota}}<br>{{Flag|South Dakota}}<br>{{Flag|Wyoming}})
|
A member of the family [[Trionychidae]].
|[[File:Axestemys_infernalis.jpg|center|frameless]]
|-
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''[[Banhxeochelys]]''<ref>{{Cite journal|author1=Rafaella C. Garbin |author2=Madelaine Böhme |author3=Walter G. Joyce |year=2019 |title=A new testudinoid turtle from the middle to late Eocene of Vietnam |journal=PeerJ |volume=7 |article-number=e6280 |doi=10.7717/peerj.6280 |pmid=30805245 |pmc=6383559 |doi-access=free }}</ref>
|
Gen. et sp. nov
|
Valid
|
Garbin, [[Madelaine Böhme|Böhme]] & Joyce
|
[[Eocene]] (late [[Bartonian]]–late [[Priabonian]])
Line 486 ⟶ 739:
|
A [[Crown group#Pan-group|pan]]-[[Geoemydidae|geoemydid]]. The type species is ''B. trani''.
|[[File:Banhxeochelys_carapax_collage_-_Garbin_et_al_2019.png|center|frameless]]
|-
|
''[[Duboisemys]]''<ref>{{cite journal |author1=Hans-Volker Karl |author2=Amtyaz Safi |author3=Hans-Dieter Philippen |year=2019 |title=Evidences of cheloniophagy by early hominid (''Homo erectus'') during middle of Pleistocene from beds of Trinil's layers in central Java (Indonesia), with an updated list of Trinil's Testudines, and a redescription of ''Duboisemys isoclina'' (Dubois, 1908) |journal=International Journal of Zoology Studies |volume=4 |issue=6 |pages=73–84 |url=https://www.researchgate.net/publication/338411328 }}</ref>
|
Gen. et comb. nov
|
Valid
|
Karl, Safi & Philippen
|
[[Middle Pleistocene]]
|
[[Trinil]] Beds
|
{{Flag|Indonesia}}
|
A member of the family [[Geoemydidae]]. The type species is ''"[[Hardella]]" isoclina'' Dubois (1908).
|
|-
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''[[Francemys]]''<ref>{{Cite journal|author=A. Pérez-García |year=2019 |title=The African Aptian ''Francemys gadoufaouaensis'' gen. et sp. nov.: new data on the early diversification of Pelomedusoides (Testudines, Pleurodira) in northern Gondwana |journal=Cretaceous Research |volume=102 |pages=112–126 |doi=10.1016/j.cretres.2019.06.003 |s2cid=197575106 |doi-access=free }}</ref>
|
Gen. et sp. nov
Line 507 ⟶ 778:
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''[[Ilatardia]]''<ref>{{Cite journal|author=Adán Pérez-García |year=2019 |title=A new member of Taphrosphyini (Pleurodira, Bothremydidae) from the Maastrichtian of Niger |journal=Journal of African Earth Sciences |volume=158 |pagesarticle-number=Article 103548 |doi=10.1016/j.jafrearsci.2019.103548 |bibcode=2019JAfES.15803548P |s2cid=198409262 }}</ref>
|
Gen. et sp. nov
Line 524 ⟶ 795:
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''[[Kalasinemys]]''<ref>{{Cite journal|author1=Haiyan Tong |author2=Wilailuck Naksri |author3=Eric Buffetaut |author4=Suravech Suteethorn |author5=Varavudh Suteethorn |author6=Phornphen Chantasit |author7=Julien Claude |year=2019 |title=''Kalasinemys'', a new xinjiangchelyid turtle from the Late Jurassic of NE Thailand |journal=Geological Magazine |volume=in press156 |issue=10 |pages=1645–1656 1–12|doi=10.1017/S0016756818000791 |bibcode=2019GeoM..156.1645T |s2cid=134746960 }}</ref>
|
Gen. et sp. nov
|
Valid
In press
|
Tong ''et al.''
Line 542 ⟶ 813:
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|
''[[Protoshachemys]]''<ref>{{Cite journal|author1=Haiyan Tong |author2=Eric Buffetaut |author3=Varavudh Suteethorn |author4=Suravech Suteethorn |author5=Gilles Cuny |author6=Lionel Cavin |author7=Uthumporn Deesri |author8=Jeremy E. Martin |author9=Kamonrak Wongko |author10=Wilailuck Naksri |author11=Julien Claude |year=2019 |title=Phu Din Daeng, a new Early Cretaceous vertebrate locality on the Khorat Plateau, NE Thailand |journal=Annales de Paléontologie |volume=in105 press|issue=3 |pages=223–237 |doi=10.1016/j.annpal.2019.04.004 |s2cid=189968031 }}</ref>
|
Gen. et sp. nov
|
Valid
In press
|
Tong ''et al.''
Line 560 ⟶ 831:
|-
|
''[[Saxochelys]]''<ref>{{Cite journal|author1=Tyler R. Lyson |author2=Jacob L. Sayler |author3=Walter G. Joyce |year=2019 |title=A new baenid turtle, ''Saxochelys gilberti'', gen. et sp. nov., from the uppermost Cretaceous (Maastrichtian) Hell Creek Formation: sexual dimorphism and spatial niche partitioning within the most speciose group of Late Cretaceous turtles |journal=Journal of Vertebrate Paleontology |volume=39 |issue=4 |pages=e1662428 |doi=10.1080/02724634.2019.1662428 |s2cid=208587902 |url=https://figshare.com/articles/dataset/A_new_baenid_turtle_i_Saxochelys_gilberti_i_gen_et_sp_nov_from_the_uppermost_Cretaceous_Maastrichtian_Hell_Creek_Formation_sexual_dimorphism_and_spatial_niche_partitioning_within_the_most_speciose_group_of_Late_Cretaceous_turtles/9944129 |url-access=subscription }}</ref>
''[[Wutuchelys]]''<ref>{{Cite journal|author1=Haiyan Tong |author2=Julien Claude |author3=Cheng-Sen Li |author4=Jian Yang |author5=Thierry Smith |year=2019 |title=''Wutuchelys eocenica'' n. gen. n. sp., an Eocene stem testudinoid turtle from Wutu, Shandong Province, China |journal=Geological Magazine |volume=156 |issue=1 |pages=133–146 |doi=10.1017/S0016756817000905 }}</ref>
|
Gen. et sp. nov
Line 566 ⟶ 837:
Valid
|
Lyson, Sayler & Joyce
Tong ''et al.''
|
[[Late Cretaceous]] ([[Maastrichtian]])
Early [[Eocene]]
|
[[WutuHell Creek Formation]]
|
{{Flag|United States}}<br>({{Flag|North Dakota}})
{{Flag|China}}
|
A [[Crownmember group#Stemof the family groups|stem]]-[[Testudinoidea|testudinoidBaenidae]]. Genus includes new species ''WS. eocenicagilberti''.
|
|-
|
''[[YaminuechelysTasbacka|YaminuechelysTasbacka sulcipeculiarisgermanica]]''<ref>{{Citecite journal |author1=CarolinaHans-Volker OriozabalaKarl |author2=JulianaElke SterliGröning |author3=MarceloCarsten S. De La FuenteBrauckmann |year=2019 |title=New''Tasbacka speciesgermanica'' ofn. thesp., long-neckeda chelidnew ''Yaminuechelys''Campanian frommarine theturtle Upperfrom CretaceousNW-Germany (Campanian–MaastrichtianTestudines: Chelonioidea) of Chubut, Argentina |journal=CretaceousMainzer Naturwissenschaftliches ResearchArchiv |volume=in press56 |pages= |doi=10.1016/j.cretres.2019.10419799–112 }}</ref>
|
Sp. nov
|
Valid
In press
|
Karl, Gröning & Brauckmann
Oriozabala, Sterli & De La Fuente
|
[[Late Cretaceous]] ([[Campanian]]–[[Maastrichtian]])
|
[[La Colonia Formation]]
|
{{Flag|ArgentinaGermany}}
|
|
A member of the family [[Chelidae]].
|-
|
''[[Trinitichelys|"Trinitichelys" maini]]''<ref>{{cite journal |author1=Brent Adrian |author2=Heather F. Smith |author3=Christopher R. Noto |author4=Aryeh Grossman |year=2019 |title=A new baenid, ''"Trinitichelys" maini'' sp. nov., and other fossil turtles from the Upper Cretaceous Arlington Archosaur Site (Woodbine Formation, Cenomanian), Texas, USA |journal=Palaeontologia Electronica |volume=22 |issue=3 |pages=Article number 22.3.81 |doi=10.26879/1001 |s2cid=210136178 |doi-access=free }}</ref>
|
Sp. nov
|
Valid
|
Adrian ''et al.''
|
[[Late Cretaceous]] ([[Cenomanian]])
|
[[Woodbine Formation]]
|
{{Flag|United States}}<br>({{Flag|Texas}})
|
A member of the family [[Baenidae]].
|
|-
|
''[[Wutuchelys]]''<ref>{{Cite journal|author1=Haiyan Tong |author2=Julien Claude |author3=Cheng-Sen Li |author4=Jian Yang |author5=Thierry Smith |year=2019 |title=''Wutuchelys eocenica'' n. gen. n. sp., an Eocene stem testudinoid turtle from Wutu, Shandong Province, China |journal=Geological Magazine |volume=156 |issue=1 |pages=133–146 |doi=10.1017/S0016756817000905 |bibcode=2019GeoM..156..133T |s2cid=134788782 }}</ref>
|
Gen. et sp. nov
|
Valid
|
Tong ''et al.''
|
Early [[Eocene]]
|
[[Wutu Formation]]
|
{{Flag|China}}
|
A [[Crown group#Stem groups|stem]]-[[Testudinoidea|testudinoid]]. Genus includes new species ''W. eocenica''.
|
|-
Line 600 ⟶ 905:
 
===General research===
* A study on the impact of the [[Triassic–Jurassic extinction event]] on archosauromorph reptiles is published by Allen ''et al.'' (2019).<ref>{{Cite journal|author1=Bethany J. Allen |author2=Thomas L. Stubbs |author3=Michael J. Benton |author4=Mark N. Puttick |year=2019 |title=Archosauromorph extinction selectivity during the Triassic–Jurassic mass extinction |journal=Palaeontology |volume=62 |issue=2 |pages=211–224 |doi=10.1111/pala.12399 |s2cid=55009185 |doi-access=free |hdl=1983/e3fc2e40-c849-42ed-99fe-ea17fc26b2ec |hdl-access=free }}</ref>
* A diverse assemblage of [[Archosauriformes|archosauriform]] teeth will beis described from the [[Upper Triassic]] [[Tiki Formation]] ([[India]]) by Ray, Bhat & Datta (2019).<ref>{{cite journal |author1=Sanghamitra Ray |author2=Mohd Shafi Bhat |author3=P. M. Datta |year=2019 |title=First record of varied archosauriforms from the Upper Triassic of India based on isolated teeth, and their biostratigraphic implications |journal=Historical Biology: An International Journal of Paleobiology |volume=in press33 |issue=2 |pages=237–253 1–17|doi=10.1080/08912963.2019.1609957 |issn=0891-2963 |s2cid=155986791 }}</ref>
* A study on archosauriform teeth assemblage from the [[Middle Triassic]] [[Manda Beds]] ([[Tanzania]]), aiming to determine the taxonomic composition of that collection and its implications for the knowledge of the diversity and evolution of archosauriforms from the Manda Beds, is published by Hoffman ''et al.'' (2019).<ref>{{cite journal |author1=Devin K. Hoffman |author2=Hunter R. Edwards |author3=Paul M. Barrett |author4=Sterling J. Nesbitt |year=2019 |title=Reconstructing the archosaur radiation using a Middle Triassic archosauriform tooth assemblage from Tanzania |journal=PeerJ |volume=7 |article-number=e7970 |doi=10.7717/peerj.7970 |pmid=31720109 |pmc=6839518 |doi-access=free }}</ref>
 
===Archosaurs===
Line 609 ⟶ 915:
 
====Research====
* Virtual [[endocast]] of ''[[Proterosuchus]] fergusi'' is reconstructed by Brown ''et al.'' (2019), who evaluate the implications of the endocranial anatomy of this species for the knowledge of its life habits.<ref>{{Cite journal|author1=Emily E. Brown |author2=Richard J. Butler |author3=Martín D. Ezcurra |author4=Bhart-Anjan S. Bhullar |author5=Stephan Lautenschlager |year=2019 |title=Endocranial anatomy and life habits of the Early Triassic archosauriform ''Proterosuchus fergusi'' |journal=Palaeontology |volume=63 |issue=2 |pages=255–282 |doi=10.1111/pala.12454 |s2cid=204271968 |url=http://pure-oai.bham.ac.uk/ws/files/71803006/Brown_et_al._in_press.pdf }}</ref>
* Redescription of the anatomy of the [[holotype]] specimen of ''[[Garjainia]] prima'' is published by Ezcurra ''et al.'' (2019), who consider ''Vjushkovia triplicostata'' to be a [[Synonym (taxonomy)|junior synonym]] of ''G. prima''.<ref>{{Cite journal|author1=Martín D. Ezcurra |author2=David J. Gower |author3=Andrey G. Sennikov |author4=Richard J. Butler |year=2019 |title=The osteology of the holotype of the early erythrosuchid ''Garjainia prima'' (Diapsida: Archosauromorpha) from the upper Lower Triassic of European Russia |journal=Zoological Journal of the Linnean Society |volume=185 |issue=3 |pages=717–783 |doi=10.1093/zoolinnean/zly061 }}</ref>
* A study on the skull anatomy and phylogenetictaxonomic relationshipsvalidity of ''[[Guchengosuchus]]Vjushkovia shiguaiensistriplicostata''  is published by Butler ''et al.'' (2019).<ref>{{Citecite journal |author1=Richard J. Butler |author2=MartínAndrey DG. EzcurraSennikov |author3=JunEmma M. Dunne Liu |author4=RolandMartin BD. Sookias Ezcurra |author5=CorwinBrandon P. Hedrick |author6=Susannah C. R. Maidment |author7=Luke E. Meade |author8=Thomas J. Raven |author9=David J. SullivanGower |year=2019 |title=TheCranial anatomy and phylogenetic positiontaxonomy of the erythrosuchid archosauriform ''Guchengosuchus"Vjushkovia shiguaiensistriplicostata"''  Huene, 1960, from the earliest MiddleEarly Triassic of ChinaEuropean Russia |journal=PeerJRoyal Society Open Science |volume=76 |pagesissue=e643511 |article-number=191289 |doi=10.77171098/peerjrsos.6435191289 |pmid=3080944331827861 |pmc=63857036894557 |bibcode=2019RSOS....691289B }}</ref>
* A study on the anatomy, [[ecomorphology]] and bone microstructure ofphylogenetic membersrelationships of ''[[ProterochampsiaGuchengosuchus]], and on their implications for inferring the lifestyles of these reptiles,shiguaiensis'' is published by Arcucci,Butler Previtera''et & Mancusoal.'' (2019).<ref>{{Cite journal|author1=AndreaRichard ArcucciJ. Butler |author2=ElenaMartín PreviteraD. Ezcurra |author3=AdrianaJun CLiu |author4=Roland B. MancusoSookias |author5=Corwin Sullivan |year=2019 |title=EcomorphologyThe anatomy and bonephylogenetic microstructureposition of Proterochampsiathe erythrosuchid archosauriform ''Guchengosuchus shiguaiensis'' from the Chañaresearliest FormationMiddle |journal=ActaTriassic Palaeontologicaof PolonicaChina |volumejournal=64PeerJ |issuevolume=17 |pagesarticle-number=157–170e6435 |doi=10.42027717/apppeerj.00536.20186435 |pmid=30809443 |pmc=6385703 |doi-access=free }}</ref>
* TwoA newstudy [[Rhadinosuchinae|rhadinosuchine]]on [[Proterochampsidae|proterochampsid]]the specimensanatomy areand describedphylogenetic fromrelationships theof ''[[Chañares FormationChalishevia]] ([[Argentina]])cothurnata'' is published by EzcurraButler ''et al.'' (2019).<ref>{{Cite journal|author1=MartínRichard DanielJ. EzcurraButler |author2=MaríaAndrey Belén VonG. BaczkoSennikov |author3=MaríaMartín JimenaD. TrotteynEzcurra |author4=JuliaDavid BrendaJ. DesojoGower |year=2019 |title=NewThe proterochampsidlast specimenserythrosuchid—a expand the morphological diversityrevision of the''Chalishevia rhadinosuchinescothurnata'' offrom the Chañareslate FormationMiddle (lowerTriassic Carnian,of northwesternEuropean Argentina)Russia |journal=AmeghinianaActa Palaeontologica Polonica |volume=5664 |issue=24 |pages=79–115757–774 |doi=10.57104202/AMGHapp.25.0500648.2019.3230 |s2cid=264102352 |doi-access=free |hdl=11336/123959 |hdl-access=free }}</ref>
* A study on the morphologyanatomy, [[ecomorphology]] and affinitiesbone microstructure of isolatedmembers of [[phytosaurProterochampsia]], teethand fromon thetheir Upperimplications Triassicfor [[Tikiinferring Formation]]the ([[India]])lifestyles willof bethese reptiles, is published by DattaArcucci, KumarPrevitera & RayMancuso (2019).<ref>{{Cite journal|author1=DebajitAndrea DattaArcucci |author2=NishantElena KumarPrevitera |author3=SanghamitraAdriana RayC. Mancuso |year=2019 |title=TaxonomicEcomorphology identificationand ofbone isolatedmicrostructure phytosaurof (Diapsida, Archosauria) teethProterochampsia from the UpperChañares Triassic of India and their significancesFormation |journal=HistoricalActa Biology:Palaeontologica An International Journal of PaleobiologyPolonica |volume=in press64 |issue=1 |pages=157–170 1–11|doi=10.10804202/08912963app.201900536.16136522018 |s2cid=134821780 |doi-access=free |hdl=11336/126031 |hdl-access=free }}</ref>
* Two new [[Rhadinosuchinae|rhadinosuchine]] [[Proterochampsidae|proterochampsid]] specimens are described from the [[Chañares Formation]] ([[Argentina]]) by Ezcurra ''et al.'' (2019).<ref>{{Cite journal|author1=Martín Daniel Ezcurra |author2=María Belén Von Baczko |author3=María Jimena Trotteyn |author4=Julia Brenda Desojo |year=2019 |title=New proterochampsid specimens expand the morphological diversity of the rhadinosuchines of the Chañares Formation (lower Carnian, northwestern Argentina) |journal=Ameghiniana |volume=56 |issue=2 |pages=79–115 |doi=10.5710/AMGH.25.05.2019.3230 |s2cid=197550418 }}</ref>
* A study on the morphology and affinities of isolated [[phytosaur]] teeth from the Upper Triassic [[Tiki Formation]] ([[India]]) is published online by Datta, Kumar & Ray (2019).<ref>{{Cite journal|author1=Debajit Datta |author2=Nishant Kumar |author3=Sanghamitra Ray |year=2019 |title=Taxonomic identification of isolated phytosaur (Diapsida, Archosauria) teeth from the Upper Triassic of India and their significances |journal=Historical Biology: An International Journal of Paleobiology |volume=33 |issue=2 |pages=272–282 |doi=10.1080/08912963.2019.1613652 |s2cid=181776675 }}</ref>
 
====New taxa====
Line 629 ⟶ 938:
|-
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''[[Antarctanax]]''<ref>{{cite journal |author1=Brandon R. Peecook |author2=Roger M. H. Smith |author3=Christian A. Sidor |year=2019 |title=A novel archosauromorph from Antarctica and an updated review of a high-latitude vertebrate assemblage in the wake of the end-Permian mass extinction |journal=Journal of Vertebrate Paleontology |volume=38 |issue=6 |pages=e1536664 |doi=10.1080/02724634.2018.1536664 |s2cid=92116260 }}</ref>
|
Gen. et sp. nov
Line 662 ⟶ 971:
|
A [[phytosaur]].
|
|-
|
''[[Volcanosuchus]]''<ref>{{cite journal |author1=Debajit Datta |author2=Sanghamitra Ray |author3=Saswati Bandyopadhyay |year=2019 |title=Cranial morphology of a new phytosaur (Diapsida, Archosauria) from the Upper Triassic of India: implications for phytosaur phylogeny and biostratigraphy |journal=Papers in Palaeontology |volume=7 |issue=2 |pages=675–708 |doi=10.1002/spp2.1292 |s2cid=213698017 |url=https://zenodo.org/record/4274207 }}</ref>
|
Gen. et sp. nov
|
Valid
|
Datta, Ray & Bandyopadhyay
|
[[Late Triassic]]
|
[[Tiki Formation]]
|
{{Flag|India}}
|
A [[phytosaur]]. Genus includes new species ''V. statisticae''.
|
|-
Line 668 ⟶ 995:
==Other reptiles==
===Research===
* New information on a specimen of the [[mesosaur]] species ''[[Stereosternum]] tumidum'' affected by congenital [[scoliosis]], first described by Szczygielski ''et al.'' (2017),<ref>{{cite journal |author1=Tomasz Szczygielski |author2=Dawid Surmik |author3=Agnieszka Kapuścińska |author4=Bruce M. Rothschild |year=2017 |title=The oldest record of aquatic amniote congenital scoliosis |journal=PLoSPLOS ONE |volume=12 |issue=9 |pages=e0185338 |doi=10.1371/journal.pone.0185338 |pmid=28934336 |pmc=5608408 |bibcode=2017PLoSO..1285338S |doi-access=free }}</ref> is published by Szczygielski ''et al.'' (2019).<ref>{{cite journal |author1=Tomasz Szczygielski |author2=Dawid Dróżdż |author3=Dawid Surmik |author4=Agnieszka Kapuścińska |author5=Bruce M. Rothschild |year=2019 |title=New tomographic contribution to characterizing mesosaurid congenital scoliosis |journal=PLoSPLOS ONE |volume=14 |issue=2 |pages=e0212416 |doi=10.1371/journal.pone.0212416 |pmid=30811483 |pmc=6392265 |bibcode=2019PLoSO..1412416S |doi-access=free }}</ref>
* DescriptionA ofstudy theon anatomybone of[[histology]] aand newgrowth specimenpatterns of ''[[KapesStereosternum (genus)|Kapes bentoni]]tumidum'' from theand ''[[Otter SandstoneBrazilosaurus]] of [[Devon]] ([[United Kingdom]], and a study on the phylogenetic relationships of this species,sanpauloensis'' is published by Zaher,Klein Coram''et & [[Michael Benton|Benton]]al.'' (2019).<ref>{{Cite journal|author1=MartaNicole ZaherKlein |author2=RobertAntoine A. CoramVerrière |author3=MichaelHeitor J.Sartorelli Benton|author4=Tanja Wintrich |author5=Jörg Fröbisch |year=2019 |title=TheMicroanatomy Middleand Triassicgrowth procolophonidof the mesosaurs ''KapesStereosternum bentonitumidum'': computedand tomography''Brazilosaurus ofsanpauloensis'' the(Reptilia, skull and skeletonParareptilia) |journal=Papers inFossil PalaeontologyRecord |volume=522 |issue=12 |pages=111–13891–110 |doi=10.10025194/spp2.1232fr-22-91-2019 |s2cid=207966143 |doi-access=free }}</ref>
* New information on the anatomy of ''[[Feeserpeton]] oklahomensis'' is presented by MacDougall ''et al.'' (2019).<ref>{{Cite journal|author1=Mark J. MacDougall |author2=Anika Winge |author3=Jasper Ponstein |author4=Maren Jansen |author5=Robert R. Reisz |author6=Jörg Fröbisch |year=2019 |title=New information on the early Permian lanthanosuchoid ''Feeserpeton oklahomensis'' based on computed tomography |journal=PeerJ |volume=7 |article-number=e7753 |doi=10.7717/peerj.7753 |pmid=31687269 |pmc=6825742 |doi-access=free }}</ref>
* A new skull ascribed to ''[[Procolophon]] trigoniceps'', so far representing the most complete and best preserved specimen collected at the [[Lower Triassic]] Sanga do Cabral Supersequence ([[Brazil]]), will be described by Silva-Neves, Modesto & Dias-da-Silva (2019).<ref>{{cite journal |author1=Eduardo Silva-Neves |author2=Sean Patrick Modesto |author3=Sérgio Dias-da-Silva |year=2019 |title=A new, nearly complete skull of ''Procolophon trigoniceps'' Owen, 1876 from the Sanga do Cabral Supersequence, Lower Triassic of Southern Brazil, with phylogenetic remarks |journal=Historical Biology: An International Journal of Paleobiology |volume=in press |issue= |pages= 1–9|doi=10.1080/08912963.2018.1512106 }}</ref>
* RedescriptionDescription of the [[pareiasaur]]anatomy speciesof a new specimen of ''"[[AnthodonKapes (reptile)|AnthodonKapes bentoni]]" haughtoni'' from the [[PermianOtter Sandstone]] of [[Usili FormationDevon]] ([[TanzaniaUnited Kingdom]]), and a study on the phylogenetic relationships of this species, is published by MaischZaher, Coram & Matzke[[Michael Benton|Benton]] (2019).<ref>{{Cite journal|author1=MichaelMarta WZaher |author2=Robert A. MaischCoram |author2author3=AndreasMichael TJ. MatzkeBenton |year=2019 |title=''AnthodonThe ?Middle Triassic procolophonid haughtoni''Kapes (V. HUENE, 1944), a pareiasaurid (Parareptiliabentoni'': Pareiasauria) from the Late Permian Usilicomputed Formationtomography of Kingori,the Ruhuhuskull Basin,and Tanzaniaskeleton |journal=NeuesPapers Jahrbuch für Geologie und Paläontologie -in AbhandlungenPalaeontology |volume=2915 |issue=21 |pages=197–204111–138 |doi=10.11271002/njgpaspp2.1232 |doi-access=free |hdl=1983/2019/07963dd2d71d-a439-404a-997f-758063f40678 |hdl-access=free }}</ref>
* [[X-ray diffraction]]A study ofon the skull anatomy and bonephylogenetic fragmentsrelationships of ''[[DeltavjatiaEmbrithosaurus]] vjatkensisschwarzi'' fromis thepublished Kotelnichonline vertebrateby fossilVan siteden ([[Russia]])Brandt, isAbdala published& by Ryanskaya ''et al.''Rubidge (2019).<ref>{{Cite journal|author1=A.Marc D.Johan RyanskayaVan den Brandt |author2=D.Fernando V. KiselevaAbdala |author3=O.Bruce P.Sidney Shilovsky |author4=E. S. ShagalovRubidge |year=2019 |title=XRDCranial studymorphology and phylogenetic relationships of the Middle Permian fossilpareiasaur ''Embrithosaurus schwarzi'' from the boneKaroo tissueBasin of South Africa |journal=PowderZoological Journal of the Linnean DiffractionSociety |volume=in press188 |issue=1 |pages=202–241 1–4|doi=10.10171093/zoolinnean/S0885715619000174zlz064 }}</ref>
* Redescription of the [[pareiasaur]] species ''"[[Anthodon (reptile)|Anthodon]]" haughtoni'' from the [[Permian]] [[Usili Formation]] ([[Tanzania]]) is published by Maisch & Matzke (2019).<ref>{{Cite journal|author1=Michael W. Maisch |author2=Andreas T. Matzke |year=2019 |title=''Anthodon ? haughtoni'' (V. HUENE, 1944), a pareiasaurid (Parareptilia: Pareiasauria) from the Late Permian Usili Formation of Kingori, Ruhuhu Basin, Tanzania |journal=Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen |volume=291 |issue=2 |pages=197–204 |doi=10.1127/njgpa/2019/0796 |s2cid=134218980 }}</ref>
* A study on the composition and structure of bone fragments of ''Deltavjatia vjatkensis'' from the Kotelnich vertebrate fossil site is published by Kiseleva ''et al.'' (2019), who report [[white blood cell]]-like structures, interpreted as possible leukocytes.<ref>{{Cite journal|author1=Daria Kiseleva |author2=Oleg Shilovsky |author3=Evgeny Shagalov |author4=Anastasia Ryanskaya |author5=Maria Chervyakovskaya |author6=Elizaveta Pankrushina |author7=Nadezhda Cherednichenko |year=2019 |title=Composition and structural features of two Permian parareptile (''Deltavjatia vjatkensis'', Kotelnich Site, Russia) bone fragments and their alteration during fossilisation |journal=Palaeogeography, Palaeoclimatology, Palaeoecology |volume=526 |pages=28–42 |doi=10.1016/j.palaeo.2019.04.015 }}</ref>
* A study on the [[Ontogeny|ontogeneticX-ray diffraction]] changesstudy inof long-bone andfragments ribof ''[[histologyDeltavjatia]] of vjatkensis''Deltavjatia rossica''from andthe ''Kotelnich vertebrate fossil site ([[ScutosaurusRussia]]) karpinskii'' will beis published by BoitsovaRyanskaya ''et al.'' (2019).<ref>{{Cite journal|author1=Elizaveta A. BoitsovaD. Ryanskaya |author2=PavelD. PV. SkutschasKiseleva |author3=AndreyO. GP. SennikovShilovsky |author4=Valeriy KE. Golubev |author5=Vladimir VS. Masuytin |author6=Olga A. MasuytinaShagalov |year=2019 |title=BoneXRD histologystudy of twothe pareiasaursPermian fromfossil Russiabone (''Deltavjatia rossica'' and ''Scutosaurus karpinskii'') with implications for pareiasaurian palaeobiologytissue |journal=BiologicalPowder Journal of the Linnean SocietyDiffraction |volume=in press34 |issue=S1 |pages=S14–S17 |doi=10.10931017/biolinnean/blz094S0885715619000174 |bibcode=2019PDiff..34S..14R |s2cid=107655052 }}</ref>
* A study on the composition and structure of bone fragments of ''Deltavjatia vjatkensis'' from the Kotelnich vertebrate fossil site is published by Kiseleva ''et al.'' (2019), who report [[white blood cell]]-like structures, interpreted as possible leukocytes.<ref>{{Cite journal|author1=Daria Kiseleva |author2=Oleg Shilovsky |author3=Evgeny Shagalov |author4=Anastasia Ryanskaya |author5=Maria Chervyakovskaya |author6=Elizaveta Pankrushina |author7=Nadezhda Cherednichenko |year=2019 |title=Composition and structural features of two Permian parareptile (''Deltavjatia vjatkensis'', Kotelnich Site, Russia) bone fragments and their alteration during fossilisation |journal=Palaeogeography, Palaeoclimatology, Palaeoecology |volume=526 |pages=28–42 |doi=10.1016/j.palaeo.2019.04.015 |bibcode=2019PPP...526...28K |s2cid=146631055 }}</ref>
* A study on the microstructure of limb bones, a rib fragment and [[osteoderm]]s of ''[[Provelosaurus]] americanus'' will be published by Farias, Schultz & Soares (2019).<ref>{{cite journal |author1=Brodsky Dantas Macedo Farias |author2=Cesar Leandro Schultz |author3=Marina Bento Soares |year=2019 |title=Bone microstructure of the pareiasaur ''Provelosaurus americanus'' from the Middle Permian of southern Brazil |journal=Historical Biology: An International Journal of Paleobiology |volume=in press |issue= |pages= 1–12|doi=10.1080/08912963.2019.1617288 }}</ref>
* A study on the species[[Ontogeny|ontogenetic]] richnesschanges andin morphologicallong-bone diversityand ofrib [[Parareptilia|parareptileshistology]] overof the''Deltavjatia courserossica'' ofand their''[[Scutosaurus]] evolutionary historykarpinskii'' is published by MacDougall,Boitsova Brocklehurst''et & Fröbischal.'' (2019).<ref>{{Cite journal|author1=MarkElizaveta JA. MacDougallBoitsova |author2=NeilPavel BrocklehurstP. Skutschas |author3=JörgAndrey G. Sennikov |author4=Valeriy K. Golubev |author5=Vladimir V. Masuytin |author6=Olga A. FröbischMasuytina |year=2019 |title=SpeciesBone richnesshistology andof disparitytwo ofpareiasaurs parareptilesfrom acrossRussia the(''Deltavjatia end-Permianrossica'' massand extinction''Scutosaurus karpinskii'') with implications for pareiasaurian palaeobiology |journal=ProceedingsBiological Journal of the RoyalLinnean Society B: Biological Sciences |volume=286128 |issue=18992 |pages=Article ID 20182572289–310 |doi=10.10981093/rspb.2018.2572 |pmid=30890099 |pmc=6452079biolinnean/blz094 }}</ref>
* A study testing whetheron the consistentmicrostructure evolutionaryof sizelimb increasebones, ina rib fragment and [[Captorhinidae|captorhinidsosteoderm]]s led to major re‐patterning in theirof ''[[long boneProvelosaurus]] structureamericanus'' is published online by RomanoFarias, Schultz & RubidgeSoares (2019).<ref>{{cite journal |author1=MarcoBrodsky RomanoDantas Macedo Farias |author2=BruceCesar RubidgeLeandro Schultz |author3=Marina Bento Soares |year=2019 |title=LongBone bonemicrostructure scalingof inthe Captorhinidae:pareiasaur do''Provelosaurus limbamericanus'' bonesfrom scalethe accordingMiddle toPermian elasticof similaritysouthern in sprawling basal amniotes?Brazil |journal=LethaiaHistorical Biology: An International Journal of Paleobiology |volume=5233 |issue=3 |pages=389–402328–339 |doi=10.11111080/let08912963.123192019.1617288 |s2cid=191161997 }}</ref>
* A study on the anatomyspecies of the [[mandible]]richness and onmorphological the phylogenetic relationshipsdiversity of ''[[MoradisaurusParareptilia|parareptiles]] grandis'', based on data from new fossil material fromover the uppercourse Permianof [[Moraditheir Formation]]evolutionary of [[Niger]],history is published by ModestoMacDougall, ''etBrocklehurst al.''& Fröbisch (2019).<ref>{{citeCite journal |author1=SeanMark PJ. ModestoMacDougall |author2=CourtneyNeil D. RichardsBrocklehurst |author3=OumarouJörg Ide |author4=Christian A. SidorFröbisch |year=2019 |title=TheSpecies vertebraterichness faunaand disparity of parareptiles across the Upper end-Permian ofmass Niger—X.extinction The mandible|journal=Proceedings of the captorhinidRoyal reptileSociety ''MoradisaurusB: grandis''Biological |journal=Journal of Vertebrate PaleontologySciences |volume=38286 |issue=61899 |pagesarticle-number=e153187720182572 |doi=10.10801098/02724634rspb.2018.15318772572 |pmid=30890099 |pmc=6452079 }}</ref>
* RedescriptionA ofstudy testing whether the anatomyconsistent ofevolutionary size increase in ''[[OrovenatorCaptorhinidae|captorhinids]] mayorum''led andto amajor studyre-patterning onin thetheir phylogenetic[[long relationships of thisbone]] speciesstructure is published by FordRomano & BensonRubidge (2019), who recover both ''Orovenator'' and [[Varanopidae|varanopids]] (usually regarded as [[synapsid]]s) as [[diapsid]] reptiles.<ref>{{Citecite journal |author1=DavidMarco P. FordRomano |author2=RogerBruce B. J. BensonRubidge |year=2019 |title=ALong redescriptionbone ofscaling ''Orovenatorin mayorum''Captorhinidae: (Sauropsida,do Diapsida)limb usingbones high‐resolutionscale μCT,according andto theelastic consequencessimilarity forin earlysprawling amniotebasal phylogenyamniotes? |journal=Papers in PalaeontologyLethaia |volume=552 |issue=23 |pages=197–239389–402 |doi=10.10021111/spp2let.123612319 |s2cid=134825836 }}</ref>
* A study on the early evolutionanatomy of the diel[[mandible]] activityand patternson inthe phylogenetic relationships of diapsid''[[Moradisaurus]] lineagesgrandis'', focusingbased on data from new fossil material from the commonupper ancestorPermian branch[[Moradi ofFormation]] livingof birds[[Niger]], is published by YuModesto &''et Wangal.'' (2019).<ref>{{Citecite journal |author1=YonghuaSean WuP. Modesto |author2=HaifengCourtney D. Richards |author3=Oumarou Ide |author4=Christian A. WangSidor |year=2019 |title=ConvergentThe evolutionvertebrate fauna of bird-mammalthe sharedUpper characteristicsPermian forof adaptingNiger—X. toThe nocturnality |journal=Proceedingsmandible of the Royalcaptorhinid Societyreptile B:''Moradisaurus Biologicalgrandis'' Sciences|journal=Journal of Vertebrate Paleontology |volume=28638 |issue=18976 |pages=Article ID 20182185e1531877 |doi=10.10981080/rspb02724634.2018.2185 |pmid=309638371531877 |pmcs2cid=640889091675715 }}</ref>
* Redescription of the anatomy of ''[[Orovenator]] mayorum'' and a study on the phylogenetic relationships of this species is published by Ford & Benson (2019), who recover both ''Orovenator'' and [[Varanopidae|varanopids]] (usually regarded as [[synapsid]]s) as [[diapsid]] reptiles.<ref>{{Cite journal|author1=David P. Ford |author2=Roger B. J. Benson |year=2019 |title=A redescription of ''Orovenator mayorum'' (Sauropsida, Diapsida) using high-resolution μCT, and the consequences for early amniote phylogeny |journal=Papers in Palaeontology |volume=5 |issue=2 |pages=197–239 |doi=10.1002/spp2.1236 |s2cid=92485505 |doi-access=free }}</ref>
* A study on the [[Morphology (biology)|morphological]] diversity and rates of morphological evolution of extinct and extant [[rhynchocephalia]]ns published by Herrera-Flores, Stubbs & [[Michael Benton|Benton]] (2017)<ref>{{Cite journal|author1=Jorge A. Herrera-Flores |author2=Thomas L. Stubbs |author3=Michael J. Benton |year=2017 |title=Macroevolutionary patterns in Rhynchocephalia: is the tuatara (''Sphenodon punctatus'') a living fossil? |journal=Palaeontology |volume=60 |issue=3 |pages=319–328 |doi=10.1111/pala.12284 }}</ref> is criticized by Vaux ''et al.'' (2019).<ref>{{Cite journal|author1=Felix Vaux |author2=Mary Morgan‐Richards |author3=Elizabeth E. Daly |author4=Steven A. Trewick |year=2019 |title=Tuatara and a new morphometric dataset for Rhynchocephalia: Comments on Herrera‐Flores ''et al.'' |journal=Palaeontology |volume=62 |issue=2 |pages=321–334 |doi=10.1111/pala.12402 }}</ref><ref>{{Cite journal|author1=Jorge A. Herrera-Flores |author2=Thomas L. Stubbs |author3=Michael J. Benton |year=2019 |title=Reply to comments on: Macroevolutionary patterns in Rhynchocephalia: is the tuatara (''Sphenodon punctatus'') a living fossil? |journal=Palaeontology |volume=62 |issue=2 |pages=335–338 |doi=10.1111/pala.12404 }}</ref>
* A study on the skullearly morphologyevolution of ''[[Clevosaurus]]the hudsoni''diel andactivity ''Clevosauruspatterns cambrica''in diapsid lineages, focusing on the common ancestor branch of living birds, is published by Chambi-Trowell, WhitesideYu & BentonWang (2019).<ref>{{Cite journal|author1=SofiaYonghua A.V. Chambi-TrowellWu |author2=DavidHaifeng I. Whiteside |author3=Michael J. BentonWang |year=2019 |title=DiversityConvergent inevolution rhynchocephalianof ''Clevosaurus''bird-mammal skullsshared basedcharacteristics onfor CTadapting reconstructionto ofnocturnality two|journal=Proceedings Lateof Triassicthe speciesRoyal fromSociety GreatB: BritainBiological |journal=Acta Palaeontologica PolonicaSciences |volume=64286 |issue=11897 |pagesarticle-number=41–6420182185 |doi=10.42021098/app.00569rspb.2018.2185 |pmid=30963837 |pmc=6408890 }}</ref>
* A study on the [[Morphology (biology)|morphological]] diversity and rates of morphological evolution of extinct and extant [[rhynchocephalia]]ns published by Herrera-Flores, Stubbs & [[Michael Benton|Benton]] (2017)<ref>{{Cite journal|author1=Jorge A. Herrera-Flores |author2=Thomas L. Stubbs |author3=Michael J. Benton |year=2017 |title=Macroevolutionary patterns in Rhynchocephalia: is the tuatara (''Sphenodon punctatus'') a living fossil? |journal=Palaeontology |volume=60 |issue=3 |pages=319–328 |doi=10.1111/pala.12284 |s2cid=55955230 |doi-access=free }}</ref> is criticized by Vaux ''et al.'' (2019).<ref>{{Cite journal|author1=Felix Vaux |author2=Mary Morgan-Richards |author3=Elizabeth E. Daly |author4=Steven A. Trewick |year=2019 |title=Tuatara and a new morphometric dataset for Rhynchocephalia: Comments on Herrera-Flores ''et al.'' |journal=Palaeontology |volume=62 |issue=2 |pages=321–334 |doi=10.1111/pala.12402 |s2cid=134902015 }}</ref><ref>{{Cite journal|author1=Jorge A. Herrera-Flores |author2=Thomas L. Stubbs |author3=Michael J. Benton |year=2019 |title=Reply to comments on: Macroevolutionary patterns in Rhynchocephalia: is the tuatara (''Sphenodon punctatus'') a living fossil? |journal=Palaeontology |volume=62 |issue=2 |pages=335–338 |doi=10.1111/pala.12404 |hdl=1983/846d212a-6eb6-494e-855f-e0684bede158 |s2cid=133726749 |url=https://research-information.bris.ac.uk/ws/files/198134843/Full_text_PDF_accepted_author_manuscript_.pdf }}</ref>
* A study on the skull morphology of ''[[Clevosaurus]] hudsoni'' and ''Clevosaurus cambrica'' is published by Chambi-Trowell, Whiteside & Benton (2019).<ref>{{Cite journal|author1=Sofia A.V. Chambi-Trowell |author2=David I. Whiteside |author3=Michael J. Benton |year=2019 |title=Diversity in rhynchocephalian ''Clevosaurus'' skulls based on CT reconstruction of two Late Triassic species from Great Britain |journal=Acta Palaeontologica Polonica |volume=64 |issue=1 |pages=41–64 |doi=10.4202/app.00569.2018 |s2cid=84832388 |doi-access=free }}</ref>
* A case study of an [[osteosarcoma]] affecting a [[femur]] of a specimen of ''[[Pappochelys]] rosinae'' is published by Haridy ''et al.'' (2019).<ref>{{Cite journal|author1=Yara Haridy |author2=Florian Witzmann |author3=Patrick Asbach |author4=Rainer R. Schoch |author5=Nadia Fröbisch |author6=Bruce M. Rothschild |year=2019 |title=Triassic cancer—osteosarcoma in a 240-million-year-old stem-turtle |journal=[[JAMA Oncology]] |volume=5 |issue=3 |pages=425–426 |doi=10.1001/jamaoncol.2018.6766 |pmid=30730547 |pmc=6439844 }}</ref>
* A study on the microstructure of bones of ''Pappochelys rosinae'' is published by Schoch ''et al.'' (2019).<ref>{{Cite journal|author1=Rainer R. Schoch |author2=Nicole Klein |author3=Torsten M. Scheyer |author4=Hans-Dieter Sues |year=2019 |title=Microanatomy of the stem-turtle ''Pappochelys rosinae'' indicates a predominantly fossorial mode of life and clarifies early steps in the evolution of the shell |journal=Scientific Reports |volume=9 |pagesissue=Article1 |article-number =10430 |doi=10.1038/s41598-019-46762-z |pmid=31320733 |pmc=6639533 |bibcode=2019NatSR...910430S }}</ref>
* An isolated vertebra of a [[choristodera]]n reptile is described from the [[Cenomanian]] Essen Greensand Formation ([[Germany]]) by Reiss ''et al.'' (2019), representing the first identifiable European choristoderan from the [[Kimmeridgian]]–[[Campanian]] interval reported so far.<ref>{{Cite journal|author1=Stefan Reiss |author2=Udo Scheer |author3=Sven Sachs |author4=Benjamin P. Kear |year=2019 |title=Filling the biostratigraphical gap: first choristoderan from the Lower–mid-Cretaceous interval of Europe |journal=Cretaceous Research |volume=96 |pages=135–141 |doi=10.1016/j.cretres.2018.12.009 |s2cid=134904339 }}</ref>
* Description of new fossil material of ''[[Khurendukhosaurus]]'' from the [[Albian]] [[Khuren Dukh Formation]] ([[Mongolia]]) and a study on the anatomy and phylogenetic relationships of this reptile is published by Matsumoto ''et al.'' (2019).<ref>{{Cite journal|author1=Ryoko Matsumoto |author2=Khishigjav Tsogtbaatar |author3=Shinobu Ishigaki |author4=Chinzorig Tsogtbaatar |author5=Zorig Enkhtaivan |author6=Susan E. Evans |year=2019 |title=Revealing body proportions of the enigmatic choristodere reptile ''Khurendukhosaurus'' from Mongolia |journal=Acta Palaeontologica Polonica |volume=64 |issue=2 |pages=363–377 |doi=10.4202/app.00561.2018 |s2cid=133925482 |doi-access=free }}</ref>
* ANeck studyvertebrae onof thea bonelong-necked [[histologyTanystropheidae|tanystropheid]] ofreptile ''[[Azendohsaurus]]are laaroussii'',described and on its implications forfrom the knowledge of evolution of [[endothermMiddle Triassic]]y in [[ArchosauromorphaMoenkopi Formation]], is published by CuboFormoso &''et Jalilal.'' (2019).<ref>{{Citecite journal |author1=JorgeKiersten CuboK. Formoso |author2=Nour-EddineSterling J. Nesbitt |author3=Adam C. Pritchard |author4=Michelle R. Stocker |author5=William G. JalilParker |year=2019 |title=BoneA histologylong-necked oftanystropheid ''Azendohsaurusfrom laaroussii'':the Middle Triassic Moenkopi Formation (Anisian) provides Implicationsinsights forinto the evolutionecology ofand thermometabolismbiogeography inof Archosauromorphatanystropheids |journal=PaleobiologyPalaeontologia Electronica |volume=4522 |issue=23 |pages=317–330Article number 22.3.73 |doi=10.101726879/pab.2019.13988 |s2cid=210088772 |doi-access=free |hdl=10919/99898 |hdl-access=free }}</ref>
* A study on the anatomybone of the postcranial skeleton[[histology]] of ''[[TeraterpetonAzendohsaurus]] hrynewichorumlaaroussii'', asand wellon asits onimplications for the phylogeneticknowledge relationshipsof evolution of this[[endotherm]]y species,in will[[Archosauromorpha]], beis published by PritchardCubo & [[Hans-Dieter Sues|Sues]]Jalil (2019).<ref>{{Cite journal|author1=AdamJorge C. PritchardCubo |author2=HansNour-DieterEddine SuesJalil |year=2019 |title=PostcranialBone remainshistology of ''TeraterpetonAzendohsaurus hrynewichorumlaaroussii'' (Reptilia: Archosauromorpha)Implications andfor the mosaic evolution of thethermometabolism saurianin postcranial skeletonArchosauromorpha |journal=Journal of Systematic PalaeontologyPaleobiology |volume=in press45 |issue=2 |pages=317–330 1–21|doi=10.10801017/14772019pab.20182019.13 |s2cid=155782789 |url=https://hal.sorbonne-universite.fr/hal-02969492/file/Cubo%20et%20Jalil%20-%202019%20-%20Bone%20histology%20of%20Azendohsaurus%20laaroussii%20Implic.1551249pdf }}</ref>
* PartialA [[maxilla]]study on the anatomy of athe postcranial skeleton of ''[[Hyperodapedontinae|hyperodapedontineTeraterpeton]] [[rhynchosaur]]hrynewichorum'', possessingas awell [[Morphologyas (biology)|morphology]]on thatthe differsphylogenetic from thoserelationships of other South American rhynchosaurthis species, willis bepublished describedby fromPritchard the& [[UpperHans-Dieter TriassicSues|Sues]] [[Ischigualasto Formation]] ([[Argentina]]) by Gentil & Ezcurra (2019).<ref>{{citeCite journal |author1=AdrielAdam RC. GentilPritchard |author2=MartínHans-Dieter D. EzcurraSues |year=2019 |title=APostcranial newremains rhynchosaurof maxillary''Teraterpeton toothhrynewichorum'' plate(Reptilia: morphotypeArchosauromorpha) expandsand the disparitymosaic evolution of the groupsaurian inpostcranial the Ischigualasto Formation (Late Triassic) of Northwestern Argentinaskeleton |journal=Historical Biology: An International Journal of PaleobiologySystematic Palaeontology |volume=in press17 |issue=20 |pages=1745–1765 1–8|doi=10.1080/0891296314772019.2018.14384251551249 |s2cid=91446492 }}</ref>
* APartial study[[maxilla]] onof thea anatomy[[Hyperodapedontinae|hyperodapedontine]] of[[rhynchosaur]], thepossessing braincasea and[[Morphology middle(biology)|morphology]] andthat innerdiffers earsfrom those of ''other South American rhynchosaur species, is described from the [[MesosuchusUpper Triassic]] browni''[[Ischigualasto isFormation]] published([[Argentina]]) by SobralGentil & MüllerEzcurra (2019).<ref>{{cite journal |author1=GabrielaAdriel SobralR. Gentil |author2=JohannesMartín MüllerD. Ezcurra |year=2019 |title=TheA braincasenew ofrhynchosaur ''Mesosuchusmaxillary browni''tooth (Reptilia,plate Archosauromorpha)morphotype withexpands informationthe ondisparity of the innergroup earin andthe descriptionIschigualasto ofFormation a(Late pneumaticTriassic) sinusof Northwestern Argentina |journal=PeerJHistorical Biology: An International Journal of Paleobiology |volume=731 |issue=9 |pages=e67981223–1230 |doi=10.77171080/peerj08912963.67982018.1438425 |pmids2cid=3119862090161690 |hdl=11336/93937 |pmchdl-access=6535042free }}</ref>
* A study on the anatomy of the braincase and middle and inner ears of ''[[Mesosuchus]] browni'' is published by Sobral & Müller (2019).<ref>{{cite journal |author1=Gabriela Sobral |author2=Johannes Müller |year=2019 |title=The braincase of ''Mesosuchus browni'' (Reptilia, Archosauromorpha) with information on the inner ear and description of a pneumatic sinus |journal=PeerJ |volume=7 |article-number=e6798 |doi=10.7717/peerj.6798 |pmid=31198620 |pmc=6535042 |doi-access=free }}</ref>
* A study on the anatomy of the [[holotype]] of ''[[Teyujagua]] paradoxa'' and on the phylogenetic relationships of this species is published online by Pinheiro, De Simão-Oliveira & Butler (2019).<ref>{{Cite journal|author1=Felipe L. Pinheiro |author2=Daniel De Simão-Oliveira |author3=Richard J. Butler |year=2019 |title=Osteology of the archosauromorph ''Teyujagua paradoxa'' and the early evolution of the archosauriform skull |journal=Zoological Journal of the Linnean Society |volume=189 |issue=1 |pages=378–417 |doi=10.1093/zoolinnean/zlz093 |doi-access=free }}</ref>
 
===New taxa===
Line 706 ⟶ 1,037:
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''[[Captorhinus|Captorhinus kieraniAncistronychus]]''<ref>{{Cite journal|author1=MichaelGabriel deBraga |author2=Joseph JS. BevittGonçalves |author3author2=RobertChristian RA. ReiszSidor |year=2019 |title=A new captorhiniddrepanosauromorph, from''Ancistronychus theparadoxus'' Permiann. cavegen. systemet near Richards Spursp., Oklahoma, andfrom the taxicChinle diversityFormation of ''Captorhinus''Petrified atForest thisNational localityPark, Arizona, USA |journal=Frontiers in Earth SciencePaleoBios |volume=736 |pages=Article 112ucmp_paleobios_46203 |doiurl=10.3389https:/feart/escholarship.2019.00112org/uc/item/8x7767f8 }}</ref>
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Gen. et sp. nov
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Valid
|
Gonçalves & [[Christian Sidor|Sidor]]
|
[[Late Triassic]]
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[[Chinle Formation]]
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{{Flag|United States}}<br>({{Flag|Arizona}})
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A member of [[Drepanosauromorpha]]. The type species is ''A. paradoxus''.
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''[[Captorhinus|Captorhinus kierani]]''<ref>{{Cite journal|author1=Michael deBraga |author2=Joseph J. Bevitt |author3=Robert R. Reisz |year=2019 |title=A new captorhinid from the Permian cave system near Richards Spur, Oklahoma, and the taxic diversity of ''Captorhinus'' at this locality |journal=Frontiers in Earth Science |volume=7 |article-number=112 |doi=10.3389/feart.2019.00112 |bibcode=2019FrEaS...7..112D |doi-access=free }}</ref>
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Sp. nov
Line 721 ⟶ 1,070:
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''[[Clevosaurus|Clevosaurus hadroprodonCarbonodraco]]''<ref>{{Citecite journal |author1=AnnieArjan S. HsiouMann |author2=RandallEmily LJ. NydamMcDaniel |author3=TiagoEmily R. SimõesMcColville |author4=FlávioHillary AC. PrettoMaddin |author5year=Silvio Onary2019 |author6title=Agustín''Carbonodraco G.lundi'' Martinelligen |author7=Alexandreet Liparini |author8=Paulo Rsp. Romo de Vivar Martínez |author9=Marina Bnov., Soaresthe |author10=Cesaroldest L.parareptile, Schultzfrom |author11=MichaelLinton, W.Ohio, Caldwell |year=2019 |title=Aand new clevosauridinsights frominto the Triassicearly (Carnian)radiation of Brazil and the rise of sphenodontians in Gondwanareptiles |journal=ScientificRoyal Society Open ReportsScience |volume=96 |pagesissue=Article11 |article-number 11821=191191 |doi=10.10381098/s41598-019-48297-9rsos.191191 |pmid=3141329431827854 |pmc=66941426894558 |bibcode=2019RSOS....691191M }}</ref>
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Gen. et sp. nov
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Valid
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Mann ''et al.''
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[[Carboniferous]] ([[Moscovian (Carboniferous)|Moscovian]])
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[[Allegheny Formation|Allegheny Group]]
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{{Flag|United States}}
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A member of the family [[Acleistorhinidae]]. The type species is ''C. lundi''. Announced in 2019; the correction including the required [[ZooBank]] accession number was published in 2020.<ref>{{cite journal |author1=Arjan Mann |author2=Emily J. McDaniel |author3=Emily R. McColville |author4=Hillary C. Maddin |year=2020 |title=Correction to "''Carbonodraco lundi'' gen et sp. nov., the oldest parareptile, from Linton, Ohio, and new insights into the early radiation of reptiles" |journal=Royal Society Open Science |volume=7 |issue=1 |article-number=192198 |doi=10.1098/rsos.192198 |pmid=32180991 |pmc=7029946 |bibcode=2020RSOS....792198M }}</ref>
|[[File:Carbonodraco_holotype_diagram.jpg|center|frameless]]
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''[[Clevosaurus|Clevosaurus hadroprodon]]''<ref>{{Cite journal|author1=Annie S. Hsiou |author2=Randall L. Nydam |author3=Tiago R. Simões |author4=Flávio A. Pretto |author5=Silvio Onary |author6=Agustín G. Martinelli |author7=Alexandre Liparini |author8=Paulo R. Romo de Vivar Martínez |author9=Marina B. Soares |author10=Cesar L. Schultz |author11=Michael W. Caldwell |year=2019 |title=A new clevosaurid from the Triassic (Carnian) of Brazil and the rise of sphenodontians in Gondwana |journal=Scientific Reports |volume=9 |issue=1 |article-number=11821 |doi=10.1038/s41598-019-48297-9 |pmid=31413294 |pmc=6694142 |bibcode=2019NatSR...911821H }}</ref>
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Sp. nov
Line 734 ⟶ 1,101:
{{Flag|Brazil}}
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|[[File:Clevosaurus_restoration.jpg|center|frameless]]
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''[[Coeruleodraco]]''<ref>{{Cite journal|author1=Ryoko Matsumoto |author2=Liping Dong |author3=Yuan Wang |author4=Susan E. Evans |year=2019 |title=The first record of a nearly complete choristodere (Reptilia: Diapsida) from the Upper Jurassic of Hebei Province, People's Republic of China |journal=Journal of Systematic Palaeontology |volume=17 |issue=12 |pages=1031–1048 |doi=10.1080/14772019.2018.1494220 |s2cid=92421503 |url=https://discovery.ucl.ac.uk/id/eprint/10067787/ }}</ref>
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Gen. et sp. nov
Line 752 ⟶ 1,119:
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A member of [[Choristodera]]. Genus includes new species ''C. jurassicus''.
|[[File:Coeruleodraco_paratype.jpg|center|frameless]]
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''[[Patagosphenos]]''<ref>{{Cite journal|author1=Adriel R. Gentil |author2=Federico L. Agnolin |author3=Jordi A. Garcia marsà |author4=Matias J. Motta |author5=Fernando E. Novas |year=2019 |title=Bridging the gap: sphenodont remains from the Turonian (Upper Cretaceous) of Patagonia. Palaeobiological inferences |journal=Cretaceous Research |volume=98 |pages=72–83 |doi=10.1016/j.cretres.2019.01.016 |s2cid=135429146 }}</ref>
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Gen. et sp. nov
Line 770 ⟶ 1,137:
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An [[Opisthodontia (reptile)|eilenodontine]] [[rhynchocephalia]]n. Genus includes new species ''P. watuku''.
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''[[Sclerostropheus]]''<ref>{{cite journal |author1=Stephan N.F. Spiekman |author2=Torsten M. Scheyer |year=2019 |title=A taxonomic revision of the genus ''Tanystropheus'' (Archosauromorpha, Tanystropheidae) |journal=Palaeontologia Electronica |volume=22 |issue=3 |pages=Article number 22.3.80 |doi=10.26879/1038 |s2cid=211105850 |doi-access=free }}</ref>
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Gen. et comb. nov
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Valid
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Spiekman & Scheyer
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[[Late Triassic]] ([[Norian]])
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{{Flag|Italy}}
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A member of the family [[Tanystropheidae]]; a new genus for ''"[[Tanystropheus]]" fossai'' Wild (1980).
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''[[Shihtienfenia|Shihtienfenia completus]]''<ref>{{Cite journal|author1=Jun-you Wang |author2=Jian Yi |author3=Jun Liu |year=2019 |title=The first complete pareiasaur skull from China |journal=Acta Palaeontologica Sinica |volume=58 |issue=2 |pages=216–221 |url=https://www.researchgate.net/publication/337946326 |doi=10.19800/j.cnki.aps.2019.02.007 }}</ref>
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Sp. nov
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Valid
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Wang, Yi & Liu
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[[Permian]]
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Sunjiagou Formation
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{{Flag|China}}
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A [[pareiasaur]].
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Line 777 ⟶ 1,179:
Research concerning more than one group of reptiles listed above.
 
* A revision of existing records of marine reptiles known from the Jurassic and Cretaceous of Siberia is published by Rogov ''et al.'' (2019).<ref>{{Cite journal|author1=M. A. Rogov |author2=N. G. Zverkov |author3=V. A. Zakharov |author4=M. S. Arkhangelsky |year=2019 |title=Marine reptiles and climates of the Jurassic and Cretaceous of Siberia |journal=Stratigraphy and Geological Correlation |volume=27 |issue=4 |pages=398–423 |doi=10.1134/S0869593819040051 |bibcode=2019SGC....27..398R |s2cid=201058264 }}</ref>
* Description of fossils of marine reptiles from the Late Jurassic of the Krzyżanowice locality ([[Poland]]) and a study evaluating the palaeobiogeographic implications of these fossils is published by Tyborowski & Błażejowski (2019).<ref>{{Cite journal|author1=Daniel Tyborowski |author2=Błażej Błażejowski |year=2019 |title=New marine reptile fossils from the Late Jurassic of Poland with implications for vertebrate faunas palaeobiogeography |journal=Proceedings of the Geologists' Association |volume=130 |issue=6 |pages=741–751 |doi=10.1016/j.pgeola.2019.09.004 |s2cid=210298782 }}</ref>
 
==References==
Line 783 ⟶ 1,186:
 
[[Category:2019 in paleontology]]
[[Category:2010s in paleontology]]
[[Category:2019 in science]]
[[Category:2019-related lists]]