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<math> w_k (X) = ( \sum_Y p_{X \rightarrow Y} \centerdot w_i (Y)) \centerdot ( \sum_Z p_{X \rightarrow Z} \centerdot w_j (Z)) </math>
where <math> Y </math> and <math> Z </math> are also DNA bases. <math> p_{ X\rightarrow Y} </math> is the transition probability from nucleotide <math>X</math> to nucleotide <math> Y </math> (idem for <math> p_{X \rightarrow Z} </math>). <math> w_i(Y) </math> is the partial likelihood of the daughter node <math>
i
</math>, evaluated on nucleotide <math> Y </math> (idem for <math>
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