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{{Short description|Genus of mammals, the goats}}
{{Automatic taxobox
| fossil_range = {{Fossil range|2.58|0}} Early [[Pleistocene]] – [[Holocene|Present]]
| image = Козе изнад Ст.Леонарда, Јужни Тирол, Италија.jpg
| image_caption = ''Capra'', [[St. Leonhard in Passeier]], [[Italy]]
| taxon = Capra
| authority = [[Carl Linnaeus|Linnaeus]], [[10th edition of Systema Naturae|1758]]
| type_species = ''[[Goat|Capra hircus]]''
| type_species_authority = [[Carl Linnaeus|Linnaeus]], [[10th edition of Systema Naturae|1758]]
| subdivision_ranks = Species
| subdivision = See [[#Species and subspecies|text]].
| range_map = Capra range map2.jpg
| range_map_caption = Approximate range of the ''Capra'' species
}}
'''''Capra''''' is a [[genus]] of [[mammals]], the '''goats''', comprising ten [[species]], including the [[markhor]] and several species known as [[ibex]]es. The [[Goat|domestic goat]] (''Capra hircus'') is a [[domesticated]] [[species]] derived from the [[bezoar ibex]] (''Capra aegagrus aegagrus''). It is one of the oldest domesticated species of animal—according to archaeological evidence its earliest domestication occurred in Iran at 10,000 [[Radiocarbon calibration|calibrated]] calendar years ago.<ref>{{Cite journal|title=The Initial Domestication of Goats (Capra hircus) in the Zagros Mountains 10,000 Years Ago|url=https://www.science.org/doi/10.1126/science.287.5461.2254|journal=Science|bibcode=2000Sci...287.2254Z|last1=Zeder|first1=Melinda A.|last2=Hesse|first2=Brian|year=2000|volume=287|issue=5461|pages=2254–7|doi=10.1126/science.287.5461.2254|pmid=10731145|access-date=June 30, 2022|archive-date=July 7, 2022|archive-url=https://web.archive.org/web/20220707013603/https://www.science.org/doi/10.1126/science.287.5461.2254|url-status=live|url-access=subscription}}</ref>
Wild goats are animals of [[mountain]] habitats. They are very agile and hardy, able to climb on bare rock and survive on sparse vegetation. They can be distinguished from the genus ''[[Ovis]]'', which includes sheep, by the presence of scent glands close to the feet, in the groin, and in front of the eyes, and the absence of other facial glands, and by the presence of a beard in some specimens, and of hairless [[callus]]es on the knees of the forelegs.<ref name=Parrini2009>{{cite journal | author = Parrini, F. | year = 2009 | title = ''Capra ibex'' (Artiodactyla: Bovidae) | journal = Mammalian Species | volume = 830 | pages = 1–12 | doi = 10.1644/830.1|display-authors=etal| doi-access = free }}</ref>
==Taxonomy==
All members of the genus ''Capra'' are [[bovid]]s (members of the family Bovidae), and more specifically caprines (subfamily [[Caprinae]]). As such they are [[ruminant]]s, meaning they chew the cud, and have four-chambered stomachs which play a vital role in digesting, regurgitating, and redigesting their food.
The genus has sometimes been taken to include ''[[Ovis]]'' (sheep) and ''Ammotragus'' ([[Barbary sheep]]),<ref>Ansell, W. F. H. 1972. Order Artiodactyla. Part 15. Pp. 1–84, in ''The mammals of Africa: An identification manual'' (J. Meester and H. W. Setzer, eds.) [issued 2 May 1972]. Smithsonian Institution Press, Washington, D.C., not continuously paginated. (quoted in {{MSW3 Grubb|id=14200767}})</ref> but these are usually regarded as distinct genera, leaving ''Capra'' for ibexes. In this smaller genus, some authors have recognized only two species, the markhor on one side and all other forms included in one species on the other side.<ref>Haltenorth, T. 1963. Klassifikation der Säugetiere: Artiodactyla I. ''Handbuch der Zoologie'', '''8'''(32):1–167 (quoted in {{MSW3 Grubb|id=14200767}})</ref> Today, nine wild species are usually accepted to which is added the domestic goat:<ref>{{cite journal|doi=10.1016/j.ympev.2006.04.002 |title=Evolutionary history of the genus Capra (Mammalia, Artiodactyla): Discordance between mitochondrial DNA and Y-chromosome phylogenies |date=2006 |last1=Pidancier |first1=Nathalie |last2=Jordan |first2=Steve |last3=Luikart |first3=Gordon |last4=Taberlet |first4=Pierre |journal=Molecular Phylogenetics and Evolution |volume=40 |issue=3 |pages=739–749 |pmid=16757184 |bibcode=2006MolPE..40..739P }}</ref>
* [[Wild goat|West Asian ibex]] also known as the wild goat (''Capra aegagrus'')
** [[Bezoar ibex]] (''Capra aegagrus aegagrus'')
** [[Sindh ibex]] (''Capra aegagrus blythi'')
* [[Domestic goat]] (''Capra hircus''; includes [[feral goat]]; domesticated from ''C. aegagrus'' and sometimes considered a subspecies of it)
* [[Siberian ibex|Asian ibex]] also known as the Siberian ibex (''Capra sibirica'')<ref>{{Cite journal |last=Sarasa |first=M. |date=2023 |title=Common names of the Asiatic ibex superspecies at a turning point in its taxonomy and management |journal=Animal Biodiversity and Conservation |language=en |pages=79–86 |doi=10.32800/abc.2023.46.0079 |s2cid=257346317 |issn=2014-928X|doi-access=free }}</ref>
* [[Markhor]] (''Capra falconeri'')
* [[West Caucasian tur]] (''Capra caucasica'')
* [[East Caucasian tur]] (''Capra cylindricornis'')
* [[Alpine ibex]] (''Capra ibex'')
* [[Iberian ibex]] also known as the Spanish ibex (''Capra pyrenaica'')<ref>{{Cite journal |last1=Sarasa |first1=Mathieu |last2=Alasaad |first2=Samer |last3=Pérez |first3=Jesús M. |date=2012 |title=Common names of species, the curious case of Capra pyrenaica and the concomitant steps towards the 'wild-to-domestic' transformation of a flagship species and its vernacular names |url=https://doi.org/10.1007/s10531-011-0172-3 |journal=Biodiversity and Conservation |language=en |volume=21 |issue=1 |pages=1–12 |doi=10.1007/s10531-011-0172-3 |bibcode=2012BiCon..21....1S |s2cid=254282784 |issn=1572-9710|url-access=subscription }}</ref>
* [[Nubian ibex]] (''Capra nubiana'')<ref>{{Cite book |last1=Mendelssohn |first1=Heinrich |title=Mammalia of Israel |last2=Yom-Tov |first2=Yoram |date=1999 |publisher=Israel Academy of Sciences and Humanities |isbn=978-965-208-145-2 |series=Fauna Palaestina |___location=Jerusalem}}</ref>
* [[Walia ibex]] (''Capra walie'')
The [[goats]] of the genus ''Capra'' have complex systematic relationships, which are still not completely resolved. Recent studies based on [[mitochondrial DNA]] suggest that the Asian ibex and the Nubian ibex represent distinct species, which are not very closely related to the physically similar Alpine ibex. The Alpine ibex forms a group with the Iberian ibex. The West Caucasian tur appears to be more closely related to the wild goat than to the East Caucasian tur. The markhor is relatively little separated from other forms—previously it had been considered to be a separate branch of the genus.<ref>{{cite journal | url=https://doi.org/10.1134%2FS1022795407020135 | doi=10.1134/S1022795407020135 | title=Phylogenetic reconstructions in the genus Capra (Bovidae, Artiodactyla) based on the mitochondrial DNA analysis | date=2007 | last1=Kazanskaya | first1=E. Y. | last2=Kuznetsova | first2=M. V. | last3=Danilkin | first3=A. A. | journal=Russian Journal of Genetics | volume=43 | issue=2 | pages=181–189 | url-access=subscription }}</ref>
The following [[cladogram]] of seven ''Capra'' species is based on 2022 mitochondrial evidence:<ref>{{cite journal |last1=Robin |first1=M. |last2=Ferrari |first2=G. |last3=Akgül |first3=G. |last4=Münger |first4=X. |last5=von Seth |first5=J. |last6=Schuenemann |first6=V. J. |last7=Dalén |first7=L. |last8=Grossen |first8=C. |year=2022 |title=Ancient mitochondrial and modern whole genomes unravel massive genetic diversity loss during near extinction of Alpine ibex |journal=Molecular Ecology |volume=31 |issue=13 |pages=3548–3565 |doi=10.1111/mec.16503 |pmid=35560856 |pmc=9328357 |bibcode=2022MolEc..31.3548R |doi-access=free}}</ref>
{{clade
|1={{clade
|1=[[Siberian ibex]]
|2={{clade
|1=[[Nubian ibex]]
|2={{clade
|1=[[Markhor]]
|2={{clade
|1={{clade
|1=[[Goat|Domestic goat]]
|2=[[Bezoar ibex]]
}}
|2={{clade
|1='''Alpine ibex'''
|2=[[Iberian ibex]]
}}
}}
}}
}}
}}
}}
Almost all wild goat species are [[allopatric]] (geographically separated)—the only geographical overlaps are the wild goat (''Capra aegagrus'') with the East Caucasian tur (''Capra cylindricornis''), and the markhor (''Capra falconeri'') with the Asian ibex (''Capra sibirica''). In both cases, the overlapping species do not usually interbreed in the wild, but in captivity, all ''Capra'' species can interbreed, producing fertile offspring.<ref>V. G. Heptner: ''Mammals of the Sowjetunion Vol. I UNGULATES''. Leiden, New York, 1989 {{ISBN|90-04-08874-1}}</ref>
<gallery class=center mode=nolines widths="200px" heights="200px">
File:Goats as weed control.jpg|Goats used for natural weed control
File:Boisseau à décor de bouquetins, sloughis et échassiers - Musée du Louvre Antiquités orientales SB 3174 ; AS 12672.jpg|Prehistoric terracotta pottery from [[Susa]] depicting an [[ibex]], {{circa|4200–3500}} BC
File:Ibex in the French Vanoise National Park.JPG|Ibex securely climbing rocky slope
File:Capra nubiana, Tierpark Hellabrunn – 03.jpg|Male Nubian ibex
File:Caprine heart.jpg|Caprine heart
</gallery>
==
{{Species table |genus= Capra |authority-name=[[Carl Linnaeus|Linnaeus]] |authority-year=[[10th edition of Systema Naturae|1758]] |species-count=ten|no-note=y|narrow-percent=75}}
{{Species table/row
|name = Asian ibex, Siberian ibex |binomial=[[Capra sibirica]]
|image = File:Steinbock-P1150170.jpg|image-size=180px |image-alt=
|authority-name = Pallas
|authority-year = 1776 |authority-not-original=
|range = [[Central Asia|Central]] & [[North Asia]], [[Afghanistan]], [[Western China|West]] and [[North China]] (mainly [[Xinjiang]]), [[North India|NW India]], SE [[Turkistan Region|Kazakhstan]], [[Kyrgyzstan]], [[Mongolia]], [[Northern Pakistan|North Pakistan]], [[Southern Russia|South Russia]], [[Tajikistan]], [[Uzbekistan|East Uzbekistan]].
|range-image = File:Capra sibirica.png
|range-image-size = 180px
|size =
|habitat =
|hunting =
|iucn-status = NT
|population =
|direction =
|subspecies = {{Collapsible list |expand=yes |title=Four subspecies |bullets=on
| ''C. s. sibirica''
| ''C. s. alaiana''
| ''C. s. hagenbecki''
| ''C. s. sakeen''
}}
}}
{{Species table/row
|name= Markhor|binomial=[[Capra falconeri]]
|image=File:Markhor Schraubenziege Capra falconeri Zoo Augsburg-02.jpg |image-size=180px |image-alt=
|authority-name=Wagner |authority-year=1839 |authority-not-original=yes
|range= [[South Asia]]; the [[Karakoram]] and [[Himalayas|Himalaya]] ranges.
|range-image=File:Capra falconeri map.gif
|range-image-size=180px
|size=
|habitat=
|hunting=
|iucn-status= NT
|population=
|direction=
|subspecies={{Collapsible list |expand=yes |title=Five subspecies |bullets=on
| ''C. f. falconeri''
| ''C. f. heptneri''
| ''C. f. megaceros''
| ''C. f. cashmiriensis''
| ''C. f. jerdoni''
}}
}}
{{Species table/row
|name= Domestic goat|binomial=[[Capra hircus]]
|image=File:Gorges du Verdon Goat-Rove-black 0253.jpg|image-size=180px |image-alt=
|authority-name=Linnaeus |authority-year=1758 |authority-not-original=
|range= [[Cosmopolitan distribution|Cosmopolitan]] distribution; [[Domestication|domesticated]].
|range-image=
|range-image-size=180px
|size=
|habitat=
|hunting=
|iucn-status= LC
|population=
|direction=
|subspecies=
}}
{{Species table/row
|name=Wild goat |binomial=[[Capra aegagrus]]
|image=File:Bezoarziege.jpg|image-size=180px |image-alt=
|authority-name=Erxleben|authority-year=1777 |authority-not-original=
|range= [[Turkey]], the [[Caucasus]] to [[Turkmenistan]], Afghanistan and Pakistan
|range-image=
|range-image-size=180px
|size=
|habitat=
|hunting=
|iucn-status= NT
|population=
|direction=
|subspecies={{Collapsible list |expand=yes |title=Four subspecies |bullets=on
|[[Bezoar ibex]] ''C. a. aegagrus''
|[[Sindh ibex]] ''C. a. blythi''
|[[Chiltan ibex]] ''C. a. chialtanensis''
|[[Turkmen wild goat]] ''C. a. turcmenica''
}}
}}
{{Species table/row
|name=East Caucasian tur |binomial=[[Capra cylindricornis]]
|image=File:Daghestanischer Tur Ostkaukasischer Steinbock Capra cylindricornis Zoo Augsburg-10 (cropped).jpg|image-size=180px |image-alt=
|authority-name=Blyth |authority-year=1841 |authority-not-original=yes
|range= [[Greater Caucasus|Greater Caucasus Mountains]].
|range-image=
|range-image-size=180px
|size=
|habitat=
|hunting=
|iucn-status= NT
|population=
|direction=
|subspecies=
}}
{{Species table/row
|name=West Caucasian tur |binomial=[[Capra caucasica]]
|image=File:Capra caucasica1.JPG|image-size=180px |image-alt=
|authority-name=Güldenstädt and Pallas |authority-year=1783 |authority-not-original=
|range= Caucasus Mountains.
|range-image=
|range-image-size=180px
|size=
|habitat=
|hunting=
|iucn-status=EN
|population=
|direction=
|subspecies=
}}
{{Species table/row
|name=Alpine ibex |binomial=[[Capra ibex]]
|image=File:SteinbockGabinten.jpg|image-size=180px |image-alt=
|authority-name=Linnaeus |authority-year=1758 |authority-not-original=
|range= [[Austria]], [[Bavaria]], [[France]], [[Italy]], [[Liechtenstein]], [[Slovenia]] & [[Switzerland]].
|range-image=File:AlpineIbex distribution.png
|range-image-size=180px
|size=
|habitat=
|hunting=
|iucn-status= LC
|population=
|direction=
|subspecies=
}}
{{Species table/row
|name= Iberian ibex|binomial=[[Capra pyrenaica]]
|image=File:Cabra montés 2.jpg|image-size=180px |image-alt=
|authority-name=Schinz |authority-year=1838 |authority-not-original=yes
|range= [[Iberian Peninsula]]: [[Andorra]], [[Pyrenees|Pyrenees Mountains]], [[Spain]] to [[Portugal]].
|range-image=
|range-image-size=180px
|size=
|habitat=
|hunting=
|iucn-status= LC
|population=
|direction=
|subspecies={{Collapsible list |expand=yes |title=Four subspecies |bullets=on
|''[[Capra pyrenaica hispanica]]''
|''[[Capra pyrenaica victoriae]]''
|†''[[Capra pyrenaica lusitanica]]''
|†''[[Capra pyrenaica pyrenaica]]''
}}
}}
{{Species table/row
|name=Nubian ibex |binomial=[[Capra nubiana]]
|image=File:Nubian Ibex in Negev.JPG|image-size=180px |image-alt=
|authority-name=F. Cuvier|authority-year= 1825 |authority-not-original=
|range= [[Egypt]], [[Eritrea]], [[Ethiopia]], [[Israel]], [[Jordan]], [[Lebanon]], [[Oman]], [[Palestine (region)|Palestine]], [[Saudi Arabia]], [[Sudan]], [[United Arab Emirates]] and [[Yemen]].<ref>{{cite iucn |last1=Ross |first1=S. |last2=Elalqamy |first2=H. |last3=Al Said |first3=T. |last4=Saltz |first4=D. |title=''Capra nubiana'' |volume=2020 |page=e.T3796A22143385 |doi=10.2305/IUCN.UK.2020-2.RLTS.T3796A22143385.en}}</ref>
|range-image=
|range-image-size=180px
|size=Males: 52–74.7 kg (115–165 lb)
Females: 25.3–32.7 kg (56–72 lb)
|habitat=Mountainous desert terrain near water sources
|hunting=[[Acacia]] and other trees, shrubs, and grasses
|iucn-status= VU
|population=4,500 <ref>{{cite iucn |last1=Ross |first1=S. |last2=Elalqamy |first2=H. |last3=Al Said |first3=T. |last4=Saltz |first4=D. |title=''Capra nubiana'' |volume=2020 |page=e.T3796A22143385 |doi=10.2305/IUCN.UK.2020-2.RLTS.T3796A22143385.en}}</ref>
|direction=
|subspecies=
}}
{{Species table/row
|name= Walia ibex|binomial=[[Capra walie]]
|image=File:Walia ibex 2.jpg|image-size=180px |image-alt=
|authority-name=Rüppell|authority-year=1835 |authority-not-original=
|range= [[Ethiopian Highlands]] & [[Simien Mountains]].
|range-image=File:Capra walie distribution.png
|range-image-size=180px
|size=
|habitat=
|hunting=
|iucn-status= VU
|population=
|direction=
|subspecies=
}}
{{Species table/end}}
==Domestication and uses==
{{main|Domestic goat}}
Along with sheep, goats were among the first [[domesticated animals]]. The domestication process started at least 10,000 years ago in what is now northern [[Iran]].<ref>{{cite journal | url=https://www.science.org/doi/abs/10.1126/science.287.5461.2254 | doi=10.1126/science.287.5461.2254 | title=The Initial Domestication of Goats ( ''Capra hircus'' ) in the Zagros Mountains 10,000 Years Ago | date=2000 | last1=Zeder | first1=Melinda A. | last2=Hesse | first2=Brian | journal=Science | volume=287 | issue=5461 | pages=2254–2257 | pmid=10731145 | bibcode=2000Sci...287.2254Z | url-access=subscription }}</ref> Easy human access to goat hair, [[meat]], and [[milk]] were the primary motivations. Goat skins were popularly used until the [[Middle Ages]] for water and [[wine bottles]] when traveling and [[camping]], and in certain regions as [[parchment]] for writing.
<gallery class=center mode=nolines widths="200px" heights="200px">
File:Goats as weed control.jpg|Goats used for natural weed control
</gallery>
==References==
{{Reflist|30em}}
==External links==
{{Commons}}
* {{Cite NIE|wstitle=Ibex |year=1905 |short=x}}
{{Artiodactyla|R.3}}
{{Taxonbar|from=Q172923}}
{{Authority control}}
{{DEFAULTSORT:Capra (Genus)}}
[[Category:Capra (genus)| ]]
[[Category:Extant Pleistocene first appearances]]
[[Category:Goats]]
[[Category:Mammal genera]]
[[Category:Mammals of Africa]]
[[Category:Mammals of Asia]]
[[Category:Animal taxa named by Carl Linnaeus]]
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