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{{Short description|System of organs used for reproduction}}
{{About|the reproductive system of all types of organisms, including humans|information specific to humans|Human reproductive system}}
{{pp-move-indef}}
{{Infobox anatomy
| Name = Reproductive system
| Latin = systemata genitalia
| Sex
Sex
Precursor =
| System =
| Artery =
| Vein =
| Nerve sex
=
| Lymph =
}}
The '''reproductive system''' of an organism, also known as the '''genital system''', is the biological system made up of all the anatomical [[sex organs|organs]] involved in [[sexual reproduction]]. Many non-living substances such as fluids, [[hormone]]s, and [[pheromone]]s are also important accessories to the reproductive system.<ref>[http://training.seer.cancer.gov/module_anatomy/unit12_1_repdt_intro.html Introduction to the Reproductive System], Epidemiology and End Results (Seer) Program. {{webarchive |url=https://web.archive.org/web/20071024151454/http://training.seer.cancer.gov/module_anatomy/unit12_1_repdt_intro.html |date=October 24, 2007 }}</ref> Unlike most [[organ system]]s, the [[sex]]es of [[sexual differentiation|differentiated]] [[species]] often have significant differences. These differences allow for a combination of genetic material between two individuals, which allows for the possibility of greater [[genes|genetic]] [[Fitness (biology)|fitness]] of the [[offspring]].<ref name=Adam>[http://www.besthealth.com/besthealth/bodyguide/reftext/html/repr_sys_fin.html Reproductive System 2001] Body Guide powered by Adam</ref>
==Animals==
{{see also|Sexual reproduction#Animals}}
{{further|Sex organ#Animals}}
In mammals, the major organs of the reproductive system include the external [[genitalia]] ([[penis]] and [[vulva]]) as well as a number of internal organs, including the [[gamete]]-producing [[gonad]]s (testicles and ovaries). Diseases of the human reproductive system are very common and widespread, particularly [[Infectious disease|communicable]] [[sexually transmitted infection]]s.<ref>[http://www.cdcnpin.org/scripts/std/std.asp#2 STD's Today] {{Webarchive|url=https://web.archive.org/web/20141025182016/http://www.cdcnpin.org/scripts/std/std.asp#2 |date=2014-10-25 }} National Prevention Network, Center for Disease Control, United States Government, retrieving 2007</ref>
Most other [[vertebrate]]s have similar reproductive systems consisting of gonads, ducts, and openings. However, there is a great diversity of physical [[adaptation]]s as well as [[Reproduction|reproductive strategies]] in every group of vertebrates.
===Vertebrates===
[[Vertebrate]]s share key elements of their reproductive systems. They all have gamete-producing organs known as [[gonad]]s. In females, these gonads are then connected by [[oviduct]]s to an opening to the outside of the body, typically the cloaca, but sometimes to a unique pore such as a [[vagina]].
==== Humans ====
{{main|Human reproductive system}}
The [[human reproductive system]] usually involves [[internal fertilization]] by [[sexual intercourse]]. During this process, the male inserts their [[erect penis]] into the [[female]]'s [[vagina]] and [[ejaculate]]s [[semen]], which contains [[sperm]]. The sperm then travels through the vagina and cervix into the [[uterus]] or [[fallopian tube]]s for fertilization of the [[ovum]]. Upon successful [[human fertilization|fertilization]] and implantation, [[gestation]] of the [[fetus]] then occurs within the female's uterus for approximately nine months, this process is known as [[pregnancy]] in humans. Gestation ends with [[childbirth]], delivery following [[Childbirth|labor]]. Labor consists of the [[muscle]]s of the uterus contracting, the [[cervical dilation|cervix dilating]], and the baby passing out the vagina (the female genital organ). Human's babies and children are nearly helpless and require high levels of [[parental care]] for many years. One important type of parental care is the use of the [[mammary gland]]s in the female [[breast]]s to [[lactation|nurse]] the baby.<ref>[http://users.rcn.com/jkimball.ma.ultranet/BiologyPages/S/Sexual_Reproduction.html Sexual Reproduction in Humans.] {{Webarchive|url=https://web.archive.org/web/20180217125604/http://users.rcn.com/jkimball.ma.ultranet/BiologyPages/S/Sexual_Reproduction.html |date=2018-02-17 }} 2006. John W. Kimball. Kimball's Biology Pages, and online textbook.</ref>
The female reproductive system has two functions: The first is to produce egg cells, and the second is to protect and nourish the offspring until birth. The male reproductive system has one function, and it is to produce and deposit sperm. Humans have a high level of [[Sexual differentiation in humans|sexual differentiation]]. In addition to differences in nearly every reproductive organ, numerous differences typically occur in [[secondary sexual characteristic]]s.
=====Male=====
{{main|Male reproductive system}}
The male reproductive system is a series of organs located outside of the body and around the pelvic region of a [[male]] that contribute towards the reproduction process. The primary direct function of the male reproductive system is to provide the male sperm for fertilization of the ovum.
The major reproductive organs of the male can be grouped into three categories. The first category is sperm production and storage. Production takes place in the [[testicle]]s, which are housed in the temperature regulating [[scrotum]], immature sperm then travel to the [[epididymis|epididymides]] for development and storage. The second category is the ejaculatory fluid-producing glands which include the [[seminal vesicles]], [[prostate]], and the [[vas deferens|vasa deferentia]]. The final category are those used for copulation, and deposition of the [[spermatozoa]] ([[sperm]]) within the male, these include the [[Human penis|penis]], [[Urethra#Male|urethra]], [[vas deferens]], and [[Cowper's gland]].
Major secondary sex characteristics include larger, more muscular stature, deepened voice, [[facial hair|facial]] and [[body hair]], broad shoulders, and development of an [[Adam's apple]]. An important sexual hormone of males is [[androgen]], and particularly [[testosterone]].
The testes release a hormone that controls the development of sperm. This hormone is also responsible for the development of physical characteristics in men such as facial hair and a deep voice.
=====Female=====
{{main|Female reproductive system}}
The human female reproductive system is a series of organs primarily located inside of the body and around the [[pelvis|pelvic]] region of a [[female]] that contribute towards the [[Human reproduction|reproductive]] process. The human female reproductive system contains three main parts: the [[vulva]], which leads to the vagina, the vaginal opening, to the uterus; the [[uterus]], which holds the developing fetus; and the [[Ovary|ovaries]], which produce the female's ova. The [[breast]]s are involved during the parenting stage of reproduction, but in most classifications they are not considered to be part of the female reproductive system.
The vagina meets the outside at the [[vulva]], which also includes the [[labia (genitalia)|labia]], [[clitoris]] and [[Urethra#Female|urethra]]; during intercourse, this area is lubricated by mucus secreted by the [[Bartholin's glands]]. The vagina is attached to the uterus through the [[cervix]], while the uterus is attached to the ovaries via the [[fallopian tubes]]. Each ovary contains hundreds of ova (singular ''[[ovum]]'').
Approximately [[Menstrual cycle|every 28 days]], the [[pituitary gland]] releases a [[hormone]] that stimulates some of the ova to develop and grow. One ovum is released and it passes through the fallopian tube into the uterus. Hormones produced by the ovaries prepare the uterus to receive the ovum. The ovum will move through her [[fallopian tubes]] and awaits the sperm for fertilization to occur. When this does not occur, i.e. no sperm for fertilization, the lining of the uterus, called the [[endometrium]], and unfertilized ova are shed each cycle through the process of [[menstruation]]. If the ovum is fertilized by sperm, it will attach to the endometrium and embryonic development will begin.
====Other mammals====
{{main|Mammalian reproductive system}}
[[File:Joey in pouch.jpg|thumb|A newborn [[Joey (marsupial)|joey]] [[suckles]] from a [[teat]] found within its mother's [[Pouch (marsupial)|pouch]]]]
[[File:Didactic model of a mammal urogenital system-FMVZ USP-17.jpeg|thumb|[[Educational toy|Didactic model]] of a mammal [[urogenital system]].]]
Most [[mammal]] reproductive systems are similar, however, there are some notable differences between the non-human mammals and humans. For instance, most male mammals have [[mammal penis|a penis]] which is [[Penile sheath|stored internally until erect]], and most have a penis bone or [[baculum]].<ref>Schultz, Nicholas G., et al. "[https://academic.oup.com/icb/article/56/4/644/2198249 The baculum was gained and lost multiple times during mammalian evolution]." Integrative and comparative biology 56.4 (2016): 644-656.</ref> Additionally, both males and females of most species do not remain continually [[sexually fertile]] as humans do and the females of most mammalian species don't grow permanent [[mammaries]] like human females do either. Like humans, most groups of mammals have descended testicles found within a scrotum, however, others have descended testicles that rest on the ventral body wall, and a few groups of mammals, such as elephants, have undescended testicles found deep within their body cavities near their [[Mammalian kidney|kidney]]s.<ref>{{cite journal |vauthors=Werdelin L, Nilsonne A |title=The evolution of the scrotum and testicular descent in mammals: a phylogenetic view |journal=J. Theor. Biol. |volume=196 |issue=1 |pages=61–72 |date=January 1999 |pmid=9892556 |doi=10.1006/jtbi.1998.0821 |bibcode=1999JThBi.196...61W }}</ref>
The [[Marsupial#Reproductive system|reproductive system of marsupial]]s is unique in that the female has two vaginae, both of which open externally through one orifice but lead to different compartments within the uterus; males usually have a [[Marsupial penis|two-pronged penis]], which corresponds to the females' two vaginae.<ref name="Tyndale-Biscoe2005">{{cite book|author=C. Hugh Tyndale-Biscoe|author-link=Cecil Hugh Tyndale-Biscoe|title=Life of Marsupials|url=https://books.google.com/books?id=KqtlPZJ9y8EC|year=2005|publisher=Csiro Publishing|isbn=978-0-643-06257-3}}</ref><ref name="Hunsaker2012">{{cite book|author=Don II Hunsaker|title=The Biology of Marsupials|url=https://books.google.com/books?id=cESCLrRJGm0C|date=2 December 2012|publisher=Elsevier Science|isbn=978-0-323-14620-3}}</ref> Marsupials typically develop their offspring in an external [[Pouch (marsupial)|pouch]] containing [[teat]]s to which their newborn young ([[Joey (marsupial)|joey]]s) attach themselves for post uterine development. Also, marsupials have a unique prepenial scrotum.<ref>{{cite book |author1=Renfree, Marilyn |author2=Tyndale-Biscoe, C. H. |title=Reproductive physiology of marsupials |publisher=Cambridge University Press |___location=Cambridge, UK |year=1987 |isbn=0-521-33792-5 |url=https://books.google.com/books?id=HpjovN0vXW4C&q=marsupial+scrotum&pg=PA129 }}</ref> The {{cvt|15|mm|in|frac=8}} long newborn joey instinctively crawls and wriggles the {{cvt|15|cm|in|frac=2}}, while clinging to fur, on the way to its mother's pouch.
In regards to males, the mammalian penis has a similar structure in reptiles and a small percentage of birds while the scrotum is only present in mammals. Regarding females, the vulva is unique to mammals with no homologue in birds, reptiles, amphibians, or fish.<ref>{{cite book|last=Trefil |first=James |title= The Encyclopedia of Science and Technology |publisher=Taylor & Francis |year=2001 |page=276 |access-date=November 10, 2023 |isbn= 978-1-13675-363-3 |url= https://books.google.com/books?id=NsbADwAAQBAJ&pg=PA276}}</ref><ref>{{cite book|last1=Armstrong |first1=David M. |last2=Fitzgerald |first2=James P. |last3=Meaney |first3=Carron A. |title= Mammals of Colorado, Second Edition |publisher=University Press of Colorado |year=2011 |page=37 |access-date=November 10, 2023 |isbn= 978-1-60732-048-7 |url= https://books.google.com/books?id=S4HgCwAAQBAJ&pg=PA37}}</ref><ref>{{cite book|last1=Cobaugh |first1=Aaron M. |last2=Stoner |first2=John B. |last3=Irwin |first3=Mark D. |title= Zookeeping:An Introduction to the Science and Technology |publisher=University of Chicago Press |year=2013 |pages=124–125 |access-date=November 10, 2023 |isbn= 978-0-22692-532-5 |url= https://books.google.com/books?id=Q7TbAgAAQBAJ&pg=PA126}}</ref> The clitoris, however, can be found in some reptiles and birds.<ref name="Kotpal">
{{harvnb|Kotpal|2010|p=394}}
</ref> In place of the uterus and vagina, non-mammal [[vertebrate]] groups have an unmodified oviduct leading directly to a [[cloaca]], which is a shared exit-hole for gametes, [[urine]], and [[feces]]. Monotremes (i.e. [[platypus]] and [[echidna]]s), a group of egg-laying mammals, also lack a uterus, vagina, and vulva, and in that respect have a reproductive system resembling that of a reptile.
=====Dogs=====
{{main|Canine reproduction}}
In domestic canines, sexual maturity (puberty) occurs between the ages of 6 and 12 months for both males and females, although this can be delayed until up to two years of age for some large breeds.
=====Horses=====
{{main|Equine anatomy#Reproductive system}}
The mare's reproductive system is responsible for controlling gestation, birth, and lactation, as well as her estrous cycle and mating behavior. The stallion's reproductive system is responsible for his sexual behavior and secondary sex characteristics (such as a large crest).
=====Even-toed ungulates=====
{{excerpt|Even-toed ungulate|Genitourinary system}}
====Birds====
{{Main|Bird anatomy#Reproductive system}}
Male and female birds have a [[cloaca]], an opening through which eggs, sperm, and wastes pass. Intercourse is performed by pressing the lips of the cloacae together, which is sometimes known as an [[intromittent organ]] which is known as a phallus that is [[analogous]] to the [[mammal penis|mammals' penis]]. The female lays [[amniote|amniotic]] eggs in which the young fetus continues to develop after it leaves the female's body. Unlike most vertebrates, female birds typically have only one functional ovary and [[oviduct]].<ref>Ritchison. BIO 554/754 Ornithology. Eastern Kentucky University.</ref> As a group, birds, like mammals, are noted for their high level of parental care.
==== Reptiles ====
{{Main|Reptiles#Reproduction}}
[[Reptiles]] are almost all [[sexual dimorphism|sexually dimorphic]], and exhibit internal fertilization through the cloaca. Some reptiles lay [[egg (biology)|egg]]s while others are ovoviviparous (animals that deliver live young). Reproductive organs are found within the cloaca of reptiles. Most male reptiles have copulatory organs, which are usually retracted or inverted and stored inside the body. In turtles and crocodilians, the male has a single median penis-like organ, while male snakes and lizards each possess a [[Hemipenis|pair of penis-like organs]].
[[File:Frog in frogspawn.jpg|thumb|right|A male [[common frog]] in nuptial colors waiting for more females to come in a mass of [[Spawn (biology)|spawn]]]]
==== Amphibians ====
{{main|Amphibian#Reproduction}}
Most [[amphibian]]s exhibit external fertilization of eggs, typically within the water, though some amphibians such as [[caecilian]]s have internal fertilization.<ref>{{cite book |last=Grzimek |first=B. |year=1974 |title=Grzimek's Animal Life Encyclopedia: Volume 5 Fishes II & Amphibians. |___location=New York |publisher=Van Nostrand Reihnhold Co. |asin=B000HHFY52|pages=301–302}}</ref> All have paired, internal gonads, connected by ducts to the cloaca.
==== Fish ====
{{main|Fish reproduction}}
[[Fish]] exhibit a wide range of different reproductive strategies. Most fish, however, are [[oviparous]] and exhibit external fertilization. In this process, females use their cloaca to release large quantities of their gametes, called [[Spawn (biology)|spawn]] into the water and one or more males release "milt", a white fluid containing many sperm over the unfertilized eggs. Other species of fish are oviparous and have internal fertilization aided by pelvic or anal [[fin]]s that are modified into an [[intromittent organ]] analogous to the human penis.<ref>[http://www.lookd.com/fish/reproduction.html Fish Reproduction]</ref> A small portion of fish species are either viviparous or [[ovoviviparous]], and are collectively known as [[livebearer]]s.<ref>[http://www.advancedaquarist.com/issues/june2002/breeder.htm Science, Biology, and Terminology of Fish reproduction: Reproductive modes and strategies-part 1] {{Webarchive|url=https://web.archive.org/web/20071108130733/http://www.advancedaquarist.com/issues/june2002/breeder.htm |date=2007-11-08 }}. 2002. MARTIN MOE. THE BREEDER'S NET Online Magazine</ref>
Fish gonads are typically pairs of either ovaries or testicles. Most fish are sexually dimorphic but some species are [[hermaphrodite|hermaphroditic]] or [[unisexual]].<ref>[http://www.buschgardens.org/infobooks/BonyFish/reproduction.html Bony Fish Reproduction] {{Webarchive|url=https://web.archive.org/web/20070927223601/http://www.buschgardens.org/infobooks/BonyFish/reproduction.html |date=2007-09-27 }} 2002. SeaWorld/Busch Gardens Animal Information Database.</ref>
===Invertebrates===
{{See also|Reproductive system of gastropods|Reproductive system of planarians}}
[[Invertebrate]]s have an extremely diverse array of reproductive systems, the only commonality may be that they all lay eggs. Also, aside from [[cephalopod]]s and [[arthropod]]s, nearly all other invertebrates exhibit [[external fertilization]].
====Cephalopods====
{{main|Cephalopod#Reproduction and life cycle}}
All [[cephalopod]]s are [[sexual dimorphism|sexually dimorphic]] and reproduce by laying [[egg (biology)|egg]]s. Most cephalopods have semi-internal fertilization, in which the male places his gametes inside the female's mantle cavity or [[pallial cavity]] to fertilize the [[Ovum|ova]] found in the female's single [[ovary]].<ref>[http://www.weichtiere.at/Mollusks/Kopffuesser/haupt.html Cephalopods.] {{webarchive|url=https://web.archive.org/web/20071020065950/http://www.weichtiere.at/Mollusks/Kopffuesser/haupt.html |date=2007-10-20 }} The Living World of Molluscs. Robert Nordsieck.</ref> Likewise, male cephalopods have only a single [[testicle]]. In the female of most cephalopods the [[nidamental gland]]s aid in development of the egg.
The "penis" in most unshelled male cephalopods ([[Coleoidea]]) is a long and muscular end of the gonoduct used to transfer spermatophores to a modified arm called a [[hectocotylus]]. That in turn is used to transfer the spermatophores to the female. In species where the hectocotylus is missing, the "penis" is long and able to extend beyond the mantle cavity and transfer the spermatophores directly to the female.
====Insects====
{{main|Insect reproductive system}}
Most [[insect]]s reproduce [[oviparous]]ly, i.e. by laying [[egg]]s. The eggs are produced by the female in a pair of [[ovaries]]. [[Sperm]], produced by the male in one [[testis]] or more commonly two, is transmitted to the female during [[mating]] by means of external [[genitalia]]. The sperm is stored within the female in one or more [[spermatheca]]e. At the time of [[fertilization]], the eggs travel along [[oviduct]]s to be fertilized by the sperm and are then expelled from the body ("laid"), in most cases via an [[ovipositor]].
====Arachnids====
{{main|Arachnids#Reproduction}}
{{see also|Opiliones penis}}
Arachnids may have one or two [[gonad]]s, which are located in the abdomen. The genital opening is usually located on the underside of the second abdominal segment. In most species, the male transfers sperm to the female in a package, or [[spermatophore]]. Complex courtship rituals have evolved in many arachnids to ensure the safe delivery of the sperm to the female.<ref name=IZ>{{cite book |author=Robert D. Barnes |year=1982 |title= Invertebrate Zoology |publisher= Holt-Saunders International |___location= Philadelphia, PA|pages= 596–604|isbn= 0-03-056747-5}}</ref>
Arachnids usually lay yolky [[egg (biology)|eggs]], which hatch into immatures that resemble adults. Scorpions, however, are either [[ovoviviparous]] or [[viviparous]], depending on species, and bear live young.
==Plants==
{{main|Plant reproductive morphology}}
{{further|Sex organ#Plants}}
Among all living organisms, [[flower]]s, which are the reproductive structures of [[flowering plant|angiosperms]], are the most varied physically and show a correspondingly great diversity in methods of reproduction.<ref name=Barr02>{{Cite journal |last=Barrett |first=S.C.H. |year=2002 |title=The evolution of plant sexual diversity |url=http://labs.eeb.utoronto.ca/barrett/pdf/schb_189.pdf |journal=Nature Reviews Genetics |volume=3 |issue=4 |pages=274–284 |doi=10.1038/nrg776 |pmid=11967552 |s2cid=7424193 |access-date=2015-06-05 |archive-date=2013-05-27 |archive-url=https://web.archive.org/web/20130527102011/http://labs.eeb.utoronto.ca/barrett/pdf/schb_189.pdf |url-status=dead }}</ref> Plants that are not flowering plants ([[green alga]]e, [[moss]]es, [[Marchantiophyta|liverwort]]s, [[hornwort]]s, [[fern]]s and [[gymnosperm]]s such as [[conifer]]s) also have complex interplays between morphological adaptation and environmental factors in their sexual reproduction. The breeding system, or how the sperm from one plant fertilizes the ovum of another, depends on the reproductive morphology, and is the single most important determinant of the genetic structure of nonclonal plant populations. [[Christian Konrad Sprengel]] (1793) studied the reproduction of flowering plants and for the first time it was understood that the [[pollination]] process involved both [[Biotic component|biotic]] and [[abiotic]] interactions.
==Fungi==
{{Main|Fungi#Reproduction}}
{{further|Sex organ#Fungi}}
Fungal reproduction is complex, reflecting the differences in lifestyles and genetic makeup within this diverse kingdom of organisms.<ref>Alexopoulos ''et al''., pp. 48–56.</ref> It is estimated that a third of all fungi reproduce using more than one method of propagation; for example, reproduction may occur in two well-differentiated stages within the [[Biological life cycle|life cycle]] of a species, the [[teleomorph]] and the [[anamorph]].<ref>Kirk ''et al''., p. 633.</ref> Environmental conditions trigger genetically determined developmental states that lead to the creation of specialized structures for sexual or asexual reproduction. These structures aid reproduction by efficiently dispersing spores or spore-containing [[propagule]]s.
==See also==
{{col div|colwidth=30em}}
*[[Human anatomy|Major systems of the human body]]
*[[Reproductive system disease]]
*[[Human sexuality]]
*[[Human sexual behavior]]
*[[Plant sexuality]]
*[[Meiosis]]
{{colend}}
==References==
{{Reflist|2}}
===Cited literature===
* {{cite book | vauthors = Alexopoulos CJ, Mims CW, Blackwell M |title=Introductory Mycology | year=1996 |publisher=John Wiley and Sons |isbn=0-471-52229-5}}
* {{cite book | vauthors = Kirk PM, Cannon PF, Minter DW, Stalpers JA |title=Dictionary of the Fungi |edition=10th |publisher=CAB International |___location=Wallingford, UK |year=2008 |isbn=978-0-85199-826-8}}
* {{cite book |last=Kotpal |first=R. L. |url=https://books.google.com/books?id=m-eQUEUjG2UC&pg=PA394 |title=Modern Text Book of Zoology: Vertebrates |publisher=Rastogi Publications |year=2010 |isbn=978-81-7133-891-7}}
==External links==
{{Commons category|Reproductive system}}
{{Wikibooks|Human Physiology|The male reproductive system}}
{{Wikibooks|Human Physiology|The female reproductive system}}
{{Wikibooks|Anatomy and Physiology of Animals|Reproductive System}}
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