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|Described as earliest described octopod, though later study shows that may not even be a mollusk.<ref>{{Cite journal |last=Whalen |first=Christopher D. |last2=Landman |first2=Neil H. |date=2022-03-08 |title=Fossil coleoid cephalopod from the Mississippian Bear Gulch Lagerstätte sheds light on early vampyropod evolution |url=https://www.nature.com/articles/s41467-022-28333-5 |journal=Nature Communications |language=en |volume=13 |issue=1 |pages=1107 |doi=10.1038/s41467-022-28333-5 |issn=2041-1723}}</ref>
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Described as the world's oldest known insect, though later study shows that may be a [[Myriapoda|myriapod]].<ref>{{Cite journal |last=Haug |first=Carolin |last2=Haug |first2=Joachim T. |date=2017-05-30 |title=The presumed oldest flying insect: more likely a myriapod? |url=https://www.ncbi.nlm.nih.gov/pmc/articles/PMC5452959/ |journal=PeerJ |volume=5 |pages=e3402 |doi=10.7717/peerj.3402 |issn=2167-8359 |pmc=5452959 |pmid=28584727}}</ref>
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|Possible stem-[[chordate]].<ref>{{Cite journal |last=MALLATT |first=JON |last2=HOLLAND |first2=NICHOLAS |date=2013-04-18 |title=Pikaia gracilens Walcott: Stem Chordate, or Already Specialized in the Cambrian? |url=https://onlinelibrary.wiley.com/doi/full/10.1002/jez.b.22500 |journal=Journal of Experimental Zoology Part B: Molecular and Developmental Evolution |volume=320 |issue=4 |pages=247–271 |doi=10.1002/jez.b.22500 |issn=1552-5007}}</ref>
'''Vertebrate characters'''
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|An early primitive shark-like [[Holocephali|holocephalian]].<ref>{{Cite journal |last=Frey |first=Linda |last2=Coates |first2=Michael I. |last3=Tietjen |first3=Kristen |last4=Rücklin |first4=Martin |last5=Klug |first5=Christian |date=2020-11-17 |title=A symmoriiform from the Late Devonian of Morocco demonstrates a derived jaw function in ancient chondrichthyans |url=https://www.nature.com/articles/s42003-020-01394-2 |journal=Communications Biology |language=en |volume=3 |issue=1 |pages=1–10 |doi=10.1038/s42003-020-01394-2 |issn=2399-3642}}</ref><ref>{{Cite journal |last=Coates |first=Michael I. |last2=Gess |first2=Robert W. |last3=Finarelli |first3=John A. |last4=Criswell |first4=Katharine E. |last5=Tietjen |first5=Kristen |date=2017 |title=A symmoriiform chondrichthyan braincase and the origin of chimaeroid fishes |url=https://www.nature.com/articles/nature20806 |journal=Nature |language=en |volume=541 |issue=7636 |pages=208–211 |doi=10.1038/nature20806 |issn=1476-4687}}</ref>
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|An early [[sawfish]]-like [[Rajiformes|skate]] relative
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