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{{Evolutionary algorithms}}
In [[genetic algorithm]]s (GA), or more general, [[evolutionary algorithm]]s (EA), a '''chromosome''' (also sometimes called a '''[[genotype]]''') is a set of parameters which define a proposed solution of the problem that the evolutionary algorithm is trying to solve. The set of all solutions, also called ''individuals'' according to the biological model, is known as the ''population''.<ref name=ga-description>{{cite web|title=Introduction to genetic algorithms: IV. Genetic Algorithm|url=http://www.obitko.com/tutorials/genetic-algorithms/ga-basic-description.php|accessdate=12 August 2015}}</ref><ref name=":0">{{Cite book |
==Chromosome design==
When creating the [[genetic representation]] of a task, it is determined which decision variables and other degrees of freedom of the task should be improved by the EA and possible additional heuristics and how the [[Genetic representation#Distinction between search space and problem space|genotype-phenotype mapping]] should look like. The design of a chromosome translates these considerations into concrete data structures for which an EA then has to be selected, configured, extended, or, in the worst case, created. Finding a suitable [[Genetic representation|representation]] of the problem ___domain for a chromosome is an important consideration, as a good representation will make the search easier by limiting the [[Genetic representation#Distinction between search space and problem space|search space]]; similarly, a poorer representation will allow a larger search space.<ref name=ga-notes>{{cite web|title=Genetic algorithms|url=http://www.cse.unsw.edu.au/~billw/cs9414/notes/ml/05ga/05ga.html|accessdate=12 August 2015|archive-date=22 October 2019|archive-url=https://web.archive.org/web/20191022162416/http://www.cse.unsw.edu.au/~billw/cs9414/notes/ml/05ga/05ga.html|url-status=dead}}</ref> In this context, suitable [[mutation (genetic algorithm)|mutation]] and [[crossover (genetic algorithm)|crossover]] [[genetic operator|operators]]<ref name=":0" /> must also be found or newly defined to fit the chosen chromosome design. An important requirement for these operators is that they not only allow all points in the search space to be reached in principle, but also make this as easy as possible.<ref>{{Cite book |last=Rothlauf |first=Franz |url=http://link.springer.com/10.1007/978-3-642-88094-0 |title=Representations for Genetic and Evolutionary Algorithms |date=2002 |publisher=Physica-Verlag HD |isbn=978-3-642-88096-4 |series=Studies in Fuzziness and Soft Computing |volume=104 |___location=Heidelberg |pages=31 |doi=10.1007/978-3-642-88094-0}}</ref><ref>{{Cite book |
The following requirements must be met by a well-suited chromosome:
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* Design of the chromosome in such a way that it covers only the search space and no additional areas. so that there is no [[Genetic representation#Redundancy|redundancy]] or only as little redundancy as possible.
* Observance of [[Causality conditions|strong causality]]: small changes in the chromosome should only lead to small changes in the phenotype.<ref>{{Cite journal |
* Designing the chromosome in such a way that it excludes prohibited regions in the search space completely or as much as possible.
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=== Chromosomes with real-valued or integer genes ===
For the processing of tasks with real-valued or mixed-integer decision variables, EAs such as the [[evolution strategy]]<ref name=":3">{{Cite book |last=Schwefel |first=Hans-Paul |url= |title=Evolution and optimum seeking |date=1995 |publisher=John Wiley & Sons |isbn=0-471-57148-2 |___location=New York |oclc=30701094}}</ref> or the real-coded GAs<ref>{{Citation |
=== Chromosomes for permutations ===
[[Combinatorial optimization|Combinatorial problems]] are mainly concerned with finding an optimal sequence of a set of elementary items. As an example, consider the problem of the [[Travelling salesman problem|traveling salesman]] who wants to visit a given number of cities exactly once on the shortest possible tour. The simplest and most obvious mapping onto a chromosome is to number the cities consecutively, to interpret a resulting sequence as [[permutation]] and to store it directly in a chromosome, where one gene corresponds to the ordinal number of a city.<ref>{{Cite book |
{| class="wikitable" style="margin-left: auto; margin-right: auto; border: none; text-align:center;"
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|| 3 || 5 || 7 || 1 || 4 || 2 || 9 || 6 || 8
|}
In addition to this encoding frequently called ''path representation'', there are several other ways of representing a permutation, for example the ''ordinal representation'' or the ''matrix representation''.<ref name=":2" /><ref>{{Cite book |last=Whitley |first=Darrell |url= |title=Evolutionary computation. Vol. 1, Basic algorithms and operators |date=2000 |publisher=Institute of Physics Pub |isbn=0-585-30560-9 |editor-last=Fogel |editor-first=David B. |___location=Bristol |pages=
=== Chromosomes for co-evolution ===
When a genetic representation contains, in addition to the decision variables, additional information that influences evolution and/or the mapping of the genotype to the phenotype and is itself subject to evolution, this is referred to as ''co-evolution''. A typical example is the [[evolution strategy]] (ES), which includes one or more mutation step sizes as strategy parameters in each chromosome.<ref name=":3" /> Another example is an additional gene to control a selection heuristic for resource allocation in a scheduling tasks.<ref name=":6">{{Cite journal |
This approach is based on the assumption that good solutions are based on an appropriate selection of strategy parameters or on control gene(s) that influences genotype-phenotype mapping. The success of the ES gives evidence to this assumption.
=== Chromosomes for complex representations ===
The chromosomes presented above are well suited for processing tasks of continuous, mixed-integer, pure-integer or combinatorial optimization. For a combination of these optimization areas, on the other hand, it becomes increasingly difficult to map them to simple strings of values, depending on the task. The following extension of the gene concept is proposed by the EA GLEAM (General Learning Evolutionary Algorithm and Method) for this purpose:<ref name=":4">{{Citation |
[[File:Genmodell Chromosombeispiel.png|thumb|212x212px|Three exemplary genes matching the adjacent gene type definitions in a chromosome organized as a list.]]
[[File:Gene model gene types.png|left|thumb|224x224px|Three exemplary genes matching the adjacent gene type definitions in a chromosome organized as a list.]]
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[[File:Genetic Program Tree.png|thumb|214x214px|Syntax tree of a formula example]]
=== Chromosomes for tree representations ===
Tree representations in a chromosome are used by [[genetic programming]], an EA type for generating [[Computer program|computer programs]] or [[Electronic circuit|circuits]].<ref name=":5" /> The trees correspond to the [[Parse tree|syntax trees]] generated by a [[compiler]] as internal representation when translating a computer program. The adjacent figure shows the syntax tree of a mathematical expression as an example. Mutation operators can rearrange, change or delete subtrees depending on the represented syntax structure. Recombination is performed by exchanging suitable subtrees.<ref>{{Cite book |
== Bibliography ==
* Thomas Bäck (1996): ''[https://books.google.com/books?id=htJHI1UrL7IC
* Wolfgang Banzhaf, P. Nordin, R. Keller, F. Francone (1998): ''Genetic Programming - An Introduction'', Morgan Kaufmann, San Francisco. {{ISBN|1-55860-510-X}}
* Kenneth A. de Jong (2006): ''Evolutionary Computation: A Unified Approach.'' MIT Press, Cambridge, MA. {{ISBN|0-262-04194-4}}
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