Oenpelli python: Difference between revisions

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| status = VU
| status_system = IUCN3.1
| status_ref = <ref name=IUCN>{{cite iucn |author=Woinarski, J. |author2=Gillespie, G. |author3=Greenlees, M. |author4=McDonald, P. |author5=Fenner, A. |year=2017 |title=''Simalia oenpelliensis '' |volume=2017 |page=e.T42494211A42494251 |doi=10.2305/IUCN.UK.2017-3.RLTS.T42494211A42494251.en |access-date=15 February 2022}}</ref>
| genus = Nyctophilopython
| parent_authority = [[Wells and Wellington affair|Wells & Wellington]], 1985
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*''Australiasis oenpelliensis'' <br />{{small|— [[Richard Walter Wells|Wells]] & [[Cliff Ross Wellington|Wellington]], 1984}}
*''Nyctophilopython oenpelliensis'' <br />{{small|— Wells & Wellington, 1985}}
*''M''[''orelia'']. ''oenpelliensis'' <br />{{small|— [[Arnold G. Kluge|Kluge]], 1993}}<ref name="McD99">{{cite book|last1=McDiarmid|author-link1=:frspecies:Roy Wallace McDiarmid|first1=R. W.|last2=Campbell|first2=J. A.|author-link2=Jonathan A. Campbell|last3=Touré|first3=T.A.|year=1999|title=Snake Species of the World: A Taxonomic and Geographic Reference|volume=. Vol. 1|publisher___location=Washington, District of Columbia: |publisher=Herpetologists' League|isbn=1-893777-01-4}}</ref>
*''Simalia oenpelliensis'' <br />{{small|— [[species:Robert Graham Reynolds|Reynolds]], [[species:Matthew L. Niemiller|Niemiller]] & [[species:Liam J. Revell|Revell]], 2014}}
*''Nawaran oenpelliensis'' <br />{{small|— Esquerré et al. 2020}}
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==Taxonomy and etymology==
The Oenpelli python was assigned to a taxonomy in 1977 by [[species:Graeme Francis Gow|Gow]], who placed it in the genus ''[[Python (genus)|Python]]''.<ref name=Gow>Gow, G. F. (1977). "A New Species of ''Python'' from Arnhem Land". ''Australian Zoologist'' '''19''': 133-139. (''Python oenpelliensis'', new species)</ref> It was then categorised by [[Harold Cogger|Cogger]] and [[species:Elizabeth E. Cameron|Cameron]] as a species of ''Morelia''.<ref name="Cogger" /> In 1984, [[Wells and Wellington affair|Wells and Wellington]] placed it into a new genus ''Nyctophilopython'', and in 2014, a work by [[species:Robert Graham Reynolds|Reynolds]], [[species:Matthew L. Niemiller|Niemiller]], and [[species:Liam J. Revell|Revell]] proposed to classify it as ''[[Simalia]]''.<ref name=Reynolds>{{cite journal|url=http://faculty.umb.edu/liam.revell/pdfs/Reynolds_etal_2014.MPE.pdf|first1=R. G.|last1=Reynolds|first2=M. L.|last2=Niemiller|first3=L. J.|last3=Revell|year=2014|title=Toward a Tree-of-Life for the boas and pythons: Multilocus species-level phylogeny with unprecedented taxon sampling|journal=[[Molecular Phylogenetics and Evolution]]|volume=71|pages=201–213|doi=10.1016/j.ympev.2013.11.011|pmid=24315866|bibcode=2014MolPE..71..201G }}</ref>
 
The [[Specific name (zoology)|specific name]], ''oenpelliensis'', is derived from the [[Type locality (biology)|type locality]], which is given as "6.5 km S.W. of [[Gunbalanya, Northern Territory|Oenpelli, Northern Territory]], Australia (12°21'S, 133°01'E)".<ref name="McD99"/><ref>Beolens, Bo; Watkins, Michael; Grayson, Michael (2011). ''The Eponym Dictionary of Reptiles''. Baltimore: Johns Hopkins University Press. xiii + 296 pp. {{ISBN|978-1-4214-0135-5}}. (''Morelia oenpelliensis'', p. 193).</ref> In 2020, the name ''Nawaran'' was erected for the genus,<ref name=Esquerre>{{cite journal |last1=Esquerré |first1=Damien |last2=Donnellan |first2=Stephen |last3=Brennan |first3=Ian G. |last4=Lemmon |first4=Alan R. |last5=Moriarty Lemmon |first5=Emily |last6=Zaher |first6=Hussam |last7=Grazziotin |first7=Felipe G. |last8=Keogh |first8=J. Scott |title=Phylogenomics, Biogeography, and Morphometrics Reveal Rapid Phenotypic Evolution in Pythons After Crossing Wallace's Line |journal=Systematic Biology |year=2020 |volume=69 |issue=6 |pages=1039–1051 |doi=10.1093/sysbio/syaa024|pmid=32208482 |doi-access=free }}</ref> overlooking the available name ''Nyctophilopython'' for the species which was immediately [[Synonym (taxonomy)|synonymised]] with the resurrected ''Nyctophilopython''.<ref name=Kaiser2020>{{cite journal |last1=Kaiser |first1=Hinrich |last2=Thomson |first2=Scott A. |last3=Shea |first3=Glenn M. |title=''Nawaran'' Esquerré, Donnellan, Brennan, Lemmon, Lemmon, Zaher, Grazziotin & Keogh, 2020 is an invalid junior synonym of ''Nyctophilopython'' Wells & Wellington, 1985 (Squamata, Pythonidae): simple priority without ''Zoobank'' pre-registration |journal=Bionomina |date=2020 |volume=20 |issue=1 |pages=47–54 |doi=10.11646/bionomina.20.1.4|s2cid=234402033 }}</ref>
 
As of September 2024, [[ITIS]] and the [[IUCN Red List]] identify the ''Simalia'' classification as valid,<ref>{{cite web |url=https://www.itis.gov/servlet/SingleRpt/SingleRpt?search_topic=TSN&search_value=1094048| title=ITIS - Report: ''Simalia oenpelliensis ''}}</ref><ref>[https://www.iucnredlist.org/species/42494211/42494251 Species. Taxonomy] {{webarchive|date=24 November 2020
|url=https://web.archive.org/web/20201124191729/https://www.iucnredlist.org/species/42494211/42494251}}</ref> while [[The Reptile Database]] uses ''Nyctophilopython''.<ref name=RDB/>
 
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A large and rare species of the monotypic genus ''Nyctophilopython'', the Oenpelli python may grow to more than {{convert|4|m|ft|abbr=on}} in total length (including tail),<ref name=ReptilesAU>{{cite web|title=Oenpelli python|url=http://reptilesofaustralia.com/snakes/pythons/moenpelli.htm#.VtPK32zTmdB|work=The Reptiles of Australia|first1=J.|last1=Fowler|first2=J.|last2=Hollister|date=11 April 2015|access-date=2016-02-29}}</ref> and one specimen in captivity is reportedly more than {{convert|5|m|ft|abbr=on}} in total length.<ref name=Got6/> The species is unusually thin in proportion to its length, relative to other pythons.<ref name=JCM/>
 
The [[Dorsum (anatomy)|dorsal]] colour pattern is dark olive-brown with darkened blotches. The belly is pale and dull, varying from cream to yellow.<ref name="NT threatened">{{cite web|url=http://www.nt.gov.au/nreta/wildlife/animals/threatened/pdf/herps/oenpelli_python_vu.pdf|title=Oenpelli python. ''Morelia oenpelliensis''|last=WoinarskWoinarski|first=J.|work=Threatened Species of the Northern Territory|publisher=Northern Territory Government|access-date=2008-10-22|url-status=dead|archive-url=https://web.archive.org/web/20080903062030/http://www.nt.gov.au/nreta/wildlife/animals/threatened/pdf/herps/oenpelli_python_vu.pdf|archive-date=2008-09-03}}</ref>
 
The Oenpelli python is able to change its [[animal colouration|skin colouration]], which tends to be lighter at night and darker in the daytime.<ref name=JCM>{{cite web|url=http://www.giantconstrictingsnakes.com/MoreliaOenpellensis.html|url-status=dead|title=The Oenpelli Python, ''Simalia oenpelliensis ''|work=Giant Constricting Snakes – The Science of Large Serpents|last=Murphy|first=J. C.|access-date=2016-02-29|archive-date=2016-03-07|archive-url=https://web.archive.org/web/20160307185012/http://www.giantconstrictingsnakes.com/MoreliaOenpellensis.html}}</ref>
 
The eggs of the Oenpelli python have been described as "huge".<ref name=Mur97/> At {{convert|110.5|by|60|mm|in|abbr=on}}, they are almost twice the size of those for the related amethystine python (''[[Amethystine python|Simalia amethystina]]''), which are reported as {{convert|70|-|98|by|45-56|mm|in|abbr=on}}.<ref name=Mur97>{{cite book|last1=Murphy|first1=J. C.|last2=Henderson|first2=R. W.|year=1997|title=Tales of Giant Snakes: A Historical Natural History of Anacondas and Pythons|publisher=Krieger Pub. Co.|isbn=0-89464-995-7|pages=[https://archive.org/details/talesofgiantsnak00murp/page/37 37, 105]|url-access=registration|url=https://archive.org/details/talesofgiantsnak00murp/page/37}}</ref>
 
==Behaviour==
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The Oenpelli python occurs in a restricted range in the Northern Territory, in the sandstone outcrops of western Arnhem Land.<ref name="NT threatened"/><ref name="Cogger" />
 
The species is found in [[habitat]] located on a sandstone [[massif]], in the regions surrounding the upper reaches of the [[Cadell River|Cadell]], [[South Alligator River|South Alligator]] and [[East Alligator River|East Alligator]] rivers. It is territorial, roaming between discrete positions, such as overhangs and caves in sandstone gorges or in a shady tree. Sightings are also reported in the region's woodland, heathland, and open rocky plains. It is noted as having an association with the [[Kombalgie]] sandstone gorges.<ref name="Cogger">{{cite book|author-link=Harold Cogger|last=Cogger|first=H. G.|year=2000|title=Reptiles and Amphibians of Australia|publisher=[[New Holland Publishers|New Holland Publishers Australia]]|isbn=9781876334338}}</ref> It is said to be associated with sandstone rock outcrops with dense vegetation.<ref name=JCM/>
 
==Conservation status==