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Scrat accidentally crashes onto Mars and unintentionally annihilates all life before returning to his ship and fleeing back to Earth.
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{{Main|List of Ice Age characters}}
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* [[Jessie J]] as [[List of Ice Age characters#Brooke|Brooke]], a ground sloth who is a resident of Geotopia and the love interest of Sid<ref name=USATBipolarLlama/><ref>{{cite tweet|last1=J|first1=Jessie|title=You 'Herd' it here first! I'm in #IceAge! Follow #CollisionCourse to see who's joining me! So excited!|user=JessieJ|number=662282787366416384|access-date=November 6, 2015|date=November 5, 2015}}</ref>
* [[Nick Offerman]] as [[List of Ice Age characters#Gavin|Gavin]], a dromaeosaur<ref name="TheHuffPostReview"/> and the father of Roger and Gertie<ref name=USATBipolarLlama/>
* [[Keke Palmer]] as [[List of Ice Age characters#Peaches|Peaches]], a young female woolly mammoth
* [[Seann William Scott]] and [[Josh Peck]] as [[Crash and Eddie (Ice Age)|Crash and Eddie]], two [[opossum]]s and Ellie's brothers
* [[Simon Pegg]] as [[List of Ice Age characters#Buck|Buck]], a one-eyed [[weasel]] and a [[dinosaur]] hunter who reunites with the Herd to warn them of the coming asteroid<ref name=Buck>{{cite tweet|last1=Pegg|first1=Simon|title=Buck's back in the herd! Follow #collisioncourse and keep an eye out for who else is joining the Ice Age family.|user=simonpegg|number=662277177623359488|access-date=November 6, 2015|date=November 5, 2015}}</ref>
* [[Wanda Sykes]] as [[List of Ice Age characters#Granny|Granny]], an aging ground sloth and Sid's grandmother<ref>{{cite tweet|last1=Sykes|first1=Wanda|title=Granny's back! Follow #CollisionCourse today to see who's joining me in #IceAge! |user=iamwandasykes|number=662516923331145728|access-date=November 6, 2015|date=November 6, 2015}}</ref>
* [[Jennifer Lopez]] as [[List of Ice Age characters#Shira|Shira]], a female saber-tooth tiger
* [[Queen Latifah]] as [[Ellie (Ice Age)|Ellie]], a female woolly mammoth – Manny's wife<ref name=USATSquirrelMacy/>
* [[Stephanie Beatriz]] as [[List of Ice Age characters#Gertie|Gertie]], a dromaeosaur<ref name="TheHuffPostReview"/> and Gavin's daughter<ref name=USATBipolarLlama/><ref>{{cite web|url=https://www.instagram.com/p/94tGYSQmiT/|title=This is #Gertie. She's a dino bird in the next #IceAge movie #CollisionCourse AND IM HER VOICE!!! also in the fam are #maxgreenfield as my bro and #nickofferman as our pops! Can't wait for you to see it: coming July 22, 2016!!|work=Instagram|access-date=April 2, 2016}}</ref>
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