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* [[Ray Romano]] as [[Manny (Ice Age)|Manny]], a [[woolly mammoth]] and the leader and co-founder of the Herd<ref name=USATSquirrelMacy>{{cite news|last1=Truitt|first1=Brian|title=Exclusive: 'Ice Age' squirrel to make Macy's debut|url=http://www.usatoday.com/story/life/movies/2015/10/22/exclusive-ice-age-scrat-balloon-macys-thanksgiving-day-parade/74366580/|access-date=October 29, 2015|work=USA Today|date=October 22, 2015}}</ref>
* [[John Leguizamo]] as [[Sid (Ice Age)|Sid]], a [[ground sloth]] and the co-founder of the Herd<ref name=USATSquirrelMacy/>
* [[Denis Leary]] as [[Diego (Ice Age)|Diego]], a [[
* [[Adam DeVine]] as [[List of Ice Age characters#Julian|Julian]], a woolly mammoth and Peaches's fiancé<ref name=USATBipolarLlama>{{cite news |last=Truitt |first=Brian |title=Jesse Tyler Ferguson is a limber llama in new 'Ice Age' |url=https://www.usatoday.com/story/life/movies/2015/11/05/jesse-tyler-ferguson-ice-age-collision-course-exclusive-character-reveal/75184240/ |access-date=November 5, 2015 |work=USA Today |date=November 5, 2015 |archive-date=September 20, 2016 |archive-url=https://web.archive.org/web/20160920142737/http://www.usatoday.com/story/life/movies/2015/11/05/jesse-tyler-ferguson-ice-age-collision-course-exclusive-character-reveal/75184240/ |url-status=live }}</ref><ref>{{cite tweet|last1=Devine|first1=Adam|title=Look at me Mom I'm a walking talking Woolly Mammoth! I'm playing Julian in the new #IceAge movie #CollisionCourse !|user=ADAMDEVINE|number=662379612563898372|access-date=November 6, 2015|date=November 5, 2015}}</ref>
* [[Jesse Tyler Ferguson]] as [[List of Ice Age characters#Shangri Llama|Shangri Llama]], a yoga-loving [[llama]] and the spiritual leader of ''Geotopia''<ref name=USATBipolarLlama/>
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