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In 2390 (around 30 years after the events of ''Killzone 3''), tensions between the Vektans and Helghast have reached a boiling point. Kellan, who let himself be captured by the Helghast as part of an investigation, is being brought across The Wall as part of a prisoner exchange for a Helghast agent named Echo. After the exchange, Kellan is ordered to return to the Helghan side of The Wall in order to retrieve classified data. Kellan recovers the classified information and makes a bold escape out of the area. Using the recovered data, Sinclair sends Kellan to the ''ISA Cassandra'', a bio-tech research station, to destroy evidence of a bio-tech weapon being created by head scientist Dr. Hillary Massar. Once on the station, Kellan encounters Echo, who escapes with Massar in tow while Kellan makes it off the station just as it falls into the sun.
Upon arriving at VSA headquarters, a massive explosion rips through the building, killing several civilians and soldiers. Through the confusion, Kellan meets up with Sinclair; just as a broadcast comes through from Vladko Tyran, leader of "The Black Hand", a paramilitary terrorist group claiming responsibility for the attack. As a result of the terrorism, all Helghast citizens residing on the Vektan side of The Wall are deported to New Helghan. Sinclair sends Kellan under-cover with the rest of the refugees in order to locate Tyran. Eventually he finds Tyran and learns that he is working for [[List of Killzone characters#Jorhan Brimve Stahl|Jorhan Brimve Stahl]] (having survived the events of ''Killzone 3''), who plans to use Massar's bio-tech weapon that has been altered to target Vektans. Shortly after, Kellan is taken captive.
Kellan is approached by Echo in his cell. He informs her that Stahl plans to use Massar's bio-tech weapon to kill Vektans and half-breeds alike, including Echo (who is half-Vektan, half-Helghast). Echo (who is revealed to be Lady Hera Visari's daughter) is displeased with Visari for being in league with Stahl and planning to ignite another war. She helps Kellan escape his cell and returns him to the Vektan side of The Wall in order to convince the VSA to stand down. Upon his return, Sinclair ignores Kellan's pleas to stand down, outraged that he is siding with Echo. Instead, Sinclair informs him that the VSA discovered massive mining spires in orbit over the planet Helghan. Kellan infiltrates the main spire and finds Massar. She states that the weapon is now with Stahl, being prepped for its final test. Sinclair orders Kellan to take Massar into custody, but Echo arrives and kills Massar. Now viewed as a traitor by Sinclair, Kellan teams up with Echo, and the two descend to Helghan to stop Stahl.
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