Talk:Evolution

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Latest comment: 18 years ago by Graft in topic Common descent

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social and religious controversies

There needs to be a description about the atheist agenda to promote atheism by evolution materialism. Especially since dawkins has labeled teaching children religion is 'child abuse' like my edit that was just undone. its clearly that such a controversial topic is not only driven by passionate religious individuals as well as atheist naturalists. I think this is evidence since gravity is considered a theory and evolution is considered a fact. Wyatt 16:24, 21 March 2007 (UTC) Reply

These issues are addressed in the FAQ and other articles.
The following discussion has been closed. Please do not modify it.
Evolution has nothing to do with religion, Atheism or Richard Dawkins except in your mind. illspirit|talk 17:05, 21 March 2007 (UTC)Reply
As I explained to Wyatt on his talk page, this is simply not relevant here. Also, please read up on what theories and facts are. (The talk archives are an excellent place to start, as is the article itself). Mikker (...) 17:24, 21 March 2007 (UTC)Reply
Evolution has nothing to do with religion? you are brilliant. feel free to delete the social and religious sections Wyatt 19:20, 21 March 2007 (UTC)Reply
I'm afraid you misunderstood, Whoutz/Wyatt. Evolution, and science in general, makes no statement about God or the validity of religion, only that there is no evidence of God and that we can develop plausible models to explain the way the universe works and how it has proceeded to this point. However, these explanations may not be consistent with the mythologies of various religions, and this may give rise to controversy. I find the suggestions you make a bit incoherent, but I don't see how claims of teaching religion to be child abuse to be relevant here. If you have some specific suggestions with citations, please feel free to bring them up here. — Knowledge Seeker 20:16, 21 March 2007 (UTC)Reply
Let's all remember to maintain civility and assume good faith.  :) Gnixon 12:26, 22 March 2007 (UTC)Reply
The issue of theory vs. fact is addressed in the FAQ for this page. I believe the relationship between atheism and evolution is discussed at Objections to evolution, but not all of that article's information will fit in the summary on this page. Gnixon 12:26, 22 March 2007 (UTC)Reply
There is no such agenda with regard to "evolution materialism", something that doesn't even exist. Information about Dawkins should go in the Richard Dawkins article or possibly the atheism article. thx1138 06:14, 5 April 2007 (UTC)Reply

Evolution as bushy, and intelligence as only one adaptation

I most recently came across this in Ben Bova's FAINT ECHOES, DISTANT STARS: THE SCIENCE AND POLITICS OF FINDING LIFE BEYOND EARTH, HarperCollins, 2004, p 247. We vaguely think of evolution as a ladder with intelligence at the top. And that's just not the case. Evolution is bushy and goes in all kinds of different directions. Other useful adaptations include sharp eyes, strong legs, a keen nose, increased wingspan, a hunting strategy of sitting and waiting and thus conserving energy, having lots of offstring, long tail feathers to attract mates, thick wooly coats for mammals in cold climates, and etc, etc.

Bova also cites Stephen Jay Gould, who takes this same general view. And here’s a website giving the transcript of a Nov. ’96 interview between Stephen and political consultant/commentator David Gergen [1] . Now, Stephen doesn’t actually use the word ‘bushy’ here, which I have heard attributed to him in other contexts. But it’s a very, very good description of what he is talking about.

I agree with Mandaclair that the Huxley graphic is great for showing a previous view of evolution and it's kind of quaint in its own way, but it is definitely not the modern view! And if you look closely at the captions, they say "Gibbon, Orang, Chimpanzee, Gorilla, Man.” Yes, these five are the currently living species of great apes (six if you wish to count the bonobo chimp as a separate species). But we are cousins!

But this same idea, slightly more sophisticated, is still in wide currency. As a young boy (I'm now 44), I remember seeing a long line of about twenty hominids, as if the whole thing is so neat and orderly. It simply is not. In fact, if we list the usual cast: Ramapithecus, Sivapithecus, Oriopithecus, Australopithecus afarensis, Australopithecus africanus, Australopithecus robustus, Australopithecus boisei, Homo habilis, Homo erectus, Homo sapien neaderthalis, all those cool cats! Well, most of these guys are our cousins, not our ancestors. (The immediate ancestor of us modern humans is Homo erectus, who is also the immediate ancestor of the neanderthals. So please note that we and the neanderthals are cousins.)

Another thing I might ask in the article is a longer, fuller explanations of L-amino acids in proteins. I take it this is the left-hand amino acids vs. the right-hand amino acids. This is a topic I find fascinating but don't know too much about. And as far as the writing style itself, sometimes a piece of wrting can include a technical description, and then a resaying of the same thing in briefer everyday language. I don't suggest this as anything mechanical and required, but rather as one more feathered arrow in your writer's quiver.

I think one of our main articles on a subject, like evolution, should be long (as long as it stays good!). One of the advantages of the Internet over a set of Encyclopedia Britannicas sitting on a shelf is that bandwidth is so much cheaper than printing! Yeah, I’ll kind of jump in the middle here. I think length in and of itself is not such a bad thing. FriendlyRiverOtter 00:52, 6 April 2007 (UTC)Reply

Thanks for your comments. If you see any possible improvements to the article, please make them! (If they're big changes, probably mention them on this page.) Gnixon 19:23, 10 April 2007 (UTC)Reply

Archival

This talk page tends to fill up quickly. Can we agree on a policy for archiving old discussions? I would suggest the following:

  • Keep any discussion with a comment less than 2 weeks old. Regularly move older ones to the archives.
  • For very long but ongoing discussions, use the hat/hab tags to hide older comments. Use the reason= parameter to explain. For example, {{hat|reason=Older comments hidden to save space. Feel free to continue the discussion below.}} produces
Older comments hidden to save space. Feel free to continue the discussion below.
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blah blah blah blah blah blah

  • When someone raises a controversial subject that is addressed in the FAQ, leave the original post, but immediately use hat/hab on the inevitable flamewar that follows. For example,
Evolution is unproven! It's a theory, not a fact! User:GenesisTellsAll
This issue is addressed in the FAQ.
The following discussion has been closed. Please do not modify it.
You're an idiot. You don't even know what those words mean. Evolution is true, no matter what a bunch of stupid religious people say. User:DarwinFishEatsYou
Screw you! Evolutionists are just atheists trying to promote their anti-faith as science. They ignore clear evidence for design. User:GenesisTellsAll
Die in a fire! To refute you, I'm going to write ten pages of text proving my point, and add ten more pages of quotes from my favorite people who agree with me. User:DarwinFishEatsYou
So am I. Ready, set, go! GenesisTellsAll

...

What do you guys think? Gnixon 16:31, 23 March 2007 (UTC)Reply

An excellent suggestion Gnixon. The warning banners and FAQ do little to stop POV pushing and vandals, so addressing it in the Talk but hiding it seems reasonable. It is difficult enough to get consensus on the topic from evolution enthusiast without wasting time addressing side issues not related to the topic. I have to admit I was initially naive to the depths of concern over creationist and ID vandalisms-I thought the editors paranoid, but was I wrong. Fill spends quite a bit of time refuting such claims from creationist and ID proponents. I am shocked as some seems less than honest (not all I should amend)which does little for their cause. GetAgrippa 18:32, 23 March 2007 (UTC)Reply
Thanks. Credit where due: the hat/hab archives were EdJohnston's idea. We could all try harder to keep our comments tightly focused and avoid starting off-topic discussions. Gnixon 02:16, 24 March 2007 (UTC)Reply
I'd propose regular archiving in a simple manner, not topic-by-topic, which is too labor-intensive to be done regularly by a human, and prone to error. It probably requires a bot to do topic-by-topic archiving without tons of work, and the available bots leave something to be desired. The hat/hab scheme for boxing up topics seems fine for questions answered in the FAQ. In general I'd suggest that this Talk page is too large when it gets over 120 kb and that the archiver should leave the most recent 80kb in place. EdJohnston 18:53, 23 March 2007 (UTC)Reply
Oh yeah. I didn't mean that each topic should be archived separately. I imagined someone glancing at the page and saying, "none of the topics above here have comments within the last two weeks, so they all get archived." As for keeping the page to 80-120 kB, I think it's better to decide a reasonable time since last comment and cut on that instead. (Of course, keeping the page small puts an upper limit on that time.) This page fills up so fast that cutting on size will often remove ongoing discussions. Editors shouldn't miss the chance to comment on recent topics just because they haven't logged onto Wikipedia in the last 3 or 4 days. Gnixon 02:16, 24 March 2007 (UTC)Reply

Topical archiving

I must have missed when it happened, but this new archiving methodology isn't very useful. I remember there used to be, at the top of the discussion page, a great reference source that had archives of discussions by topic. For example, the "Evolution is only a theory" topic, which happens over and over again, had it's own link. One could go and read it, maybe realize "oh someone's said that, and it's been set aside." Now I can't find all that stuff. Anyways, all IMHO. Orangemarlin 18:16, 26 March 2007 (UTC)Reply

Archiving by topic is really useful, but also tons of work to maintain. I suspect people switched to the simpler scheme out of laziness. Keep in mind that this talk page generates about an article's length of comments every couple weeks. See also the discussion about an "Evolution Debates" archive and its deletion as a POV fork. Gnixon 18:52, 26 March 2007 (UTC)Reply
Too bad. It was nice to refer people to old arguments. If they didn't read them, we could beat them up mercilessly. It made my days so much happier. Orangemarlin 17:33, 27 March 2007 (UTC)Reply
It would be useful if someone created such a table at the top of the page with links to discussions in the archives. Gnixon 17:33, 28 March 2007 (UTC)Reply
With User:EdJohnston's help, I think I've found the pages OM refers to. They're linked to in the 2005 archives. Gnixon 04:04, 3 April 2007 (UTC)Reply

Bot archiving

To save us effort, I've set up User:MiszaBot to automatically archive conversations older than 2 weeks. Hopefully, I got all the settings right. If it causes problems, please let me know and I'll clean things up. If anyone doesn't like this idea, please say so. Or, if you like the bot but not the settings, certainly feel free to change them yourself---it's pretty easy. Cheers, Gnixon 16:17, 3 April 2007 (UTC).Reply

Theistic Evolution

There is a need for a section about Theistic Evolution. Talk about Evolution's status in big religions such as Islam, and Christianity, and Hinduism, etc. Believe it or not, there are Muslims, Christians, and Hindus who believe in Evolution. Armyrifle 23:21, 25 March 2007 (UTC)Reply

Believe it or not, Theistic evolution.--ĶĩřβȳŤįɱéØ 23:29, 25 March 2007 (UTC)Reply
Hmm, a link somewhere in the article might be nice. i kan reed 23:38, 25 March 2007 (UTC)Reply
Please see the Social and religious controversies section. — Knowledge Seeker 02:27, 26 March 2007 (UTC)Reply
This is a science article that is not about religious beliefs at all. That some religious organizations feel the need to have a religious position on a particular aspect of science is best discussed elsewhere, and linked from the "social and religious controversies" section. thx1138 06:20, 5 April 2007 (UTC)Reply

Controversy (2)

The Social and religious controversy section is probably the most neglected one in the article, but it is one of the most important for many of the new posters on this discussion page. The section has long had "citation needed" tags. It discusses both objections to evolution and controversial social theories derived from it, but the two topics are not well-separated. The paragraphs seem to have each been developed independently and don't transition well. Can we try to improve things? Gnixon 16:09, 26 March 2007 (UTC)Reply

I attempted a major revision several days ago. I thought it would be uncontroversial since I only used the previous text and the introductions of the sub-articles, but the change was reverted by someone who preferred to discuss it here first. In response, I've created a Work in Progress page and copied my edit there. I would appreciate if people would take a look, comment at the bottom of the page, and make improvements. Thanks!! Gnixon 16:09, 26 March 2007 (UTC)Reply

Thanks to the users who made comments at the WIP page. I've recently made several edits to the Controversy section, keeping their comments in mind. Particularly, instead of trying to copy in the introductions of related articles, which made the section too long, I've simply organized the section with subsections and cut redundant material. One editor argued for cutting the "Social theories" stuff, but I've left it in for now. I hope this is satisfactory to everyone. Let's work hard to keep this section short, well-referenced, and free of both anti-evolution and anti-creationist POV. Gnixon 16:31, 28 March 2007 (UTC)Reply

Deleting the page. Gnixon 19:20, 10 April 2007 (UTC)Reply

Observation

Where are the examples of new species observed to come into existence? I've heard claims about medicines being invented by evolution and things, but I don't know of any where that evolution has actually been observed? This is different than seeing a chain of similar animals, because those animals are actually distant from each other even if they followed a similar path. All I've seen is beaks getting longer or shorter, but no real macro changes or new features. It would be nice to have some statements about it, but I may have just overlooked them. Wyatt 21:28, 26 March 2007 (UTC) Reply

This issue is addressed in the FAQ.
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This issue is addressed somewhat in the FAQ. You'll also find information in Objections to evolution since some have argued that "macroevolution" has not been observed. Gnixon 22:33, 26 March 2007 (UTC)Reply
Wyatt is right, macroevolution is the main objection in recent memory. On first thought it could be unburied from Objections and placed in its lead, which would then be replicated to Evolution. I could see some objecting to this as too specific for the lead; but to not have macroevolution in the content of the Evolution article (templates at the bottom/side don't count folks) seems to be a glaring blind spot. - RoyBoy 800 03:11, 28 March 2007 (UTC)Reply
New features have been observed, such as extra limbs and digits and such, but no new species have been observed coming into existence. The closest I could find is this: "Electric Fish in Africa Could Be Example of Evolution in Action" --Savant13 12:36, 4 April 2007 (UTC)Reply
This is also addressed in the FAQ. Gnixon 12:47, 4 April 2007 (UTC)Reply

Suggestions from Mandaclair

A number of discussions with a biology lecturer.

Definition

Proposed definition of evolution for lead. General support. Concern about "biological" qualifier. Brief discussion of strategy for addressing creationist reactions.

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Mandaclair recently made some interesting edits to the introduction. They were quickly reverted because they changed the lead significantly, adding a lot of detail, but her paragraph defining evolution seemed useful, and I wonder if we could work it in somewhere without making the lead too unwieldy:

Strictly speaking, biological evolution is the process of change over time in the heritable characteristics, or traits, of a population of organisms. Heritable traits are encoded by the genetic material of an organism (usually DNA). Evolution generally results from three processes: random mutation to genetic material, random genetic drift, and non-random natural selection within populations and species. In common vernacular, evolution is also used more generally to refer to the greater outcomes of these processes, such as the diversification of all forms of life from shared ancestors, and observable changes in the fossil record over time.

The way she enumerates three processes and separates the technical definition from the vernacular could guide the introduction and first few sections of the article, especially if we can find a way to avoid getting too technical too early. By the way, she also made several good small changes to the intro that were reverted with the others. It'd be nice if someone went through the history and copied some of the changes back in. Gnixon 16:58, 27 March 2007 (UTC)Reply

I'm not an evolutionary biologist, nor play one on TV, so it really sounds good. I don't like "biological evolution", for no other reason than I'll bet some creationist will beat up on the point that it's not really "evolution". But I could be paranoid after several months of bickering with creationists on here. Furthermore, I would like one of our more scientific types to review the sentence. Sometimes someone might simplify technology so much, that the essential meaning is lost or confused. Mandaclair is a new editor, so I'm always wary until they have gone through several rounds of discussion on these pages. But, for a first pass, I'm pretty impressed. Orangemarlin 17:32, 27 March 2007 (UTC)Reply
Hi. Thanks for listening. As I mentioned on a user talk page somewhere, there is nothing "creationist" about specifying "biological evolution" as a way of distinguishing life from other systems that evolve, such as languages, societies, or the universe as a whole. And in general, I recommend not worrying too much about what creationists will (or won't) "beat up" upon, because very little progress is to be made in those [rhetorical] dialogues, anyway. Don't write this article with "defense against creationists" in mind. The only sensible thing to do is ignore them, and write the best article you can. (Comment from User:Mandaclair.)
I very, very, very strongly agree. Gnixon 02:39, 28 March 2007 (UTC)Reply
Sounds good to me. Suggest changing "common vernacular" to "everyday speech." I will see what I can do about incorporating some of the other edits.--EveRickert 18:35, 27 March 2007 (UTC)Reply

Recommendations

A number of recommendations for the article. Few responses. Proposal of "Misconceptions" section discussed in later subsection.

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If "bold editing" was a bit easier to accomplish, I might recommend the following (Comments by User:Mandaclair):

  • strip down some of the basic genetics in the article. Keep it streamlined toward Evolution. Much of the article seems like it should be a genetics article, or population genetics article. Those topics definitely play into Evolution, but in my opinion, this article gives them too much space.
Strongly agree. Gnixon 02:38, 28 March 2007 (UTC)Reply
  • include a new section entitled "common misconceptions about evolution". These can be documented, and such a section is extremely valuable to persons approaching this subject for the first time.
What misconceptions do you have in mind? A similar section once existed, but it sort of turned into "Why Creationists' objections are wrong because they don't understand stuff." That caused lots of problems. Gnixon 02:38, 28 March 2007 (UTC)Reply
(Discussed in its own subsection below.)
  • Stabilizing selection, directional selection, and DISRUPTIVE SELECTION [left out in the current version] are the three MODES of natural selection, and they are not really correctly described here (for example: all three of them favor the "beneficial" alleles and select against "harmful" ones.) Artificial selection should not be invoked in this section -- it is trivial (arguably meaningless) in the grand scheme of things, and probably better discussed in the section about the history of Evolutionary thought, since Darwin began his treatise with an examination of artificial selection, and reasoned: if humans can produce breeds and varieties (as he called them), then why couldn't nature?
  • As mentioned, reduce the adaptationist language as much as possible. Adaptations certainly can and do occur, and they are important, but it is also very important to get across that "the race is not always to the swift, nor the battle to the strong", etc. "Fitness" has nothing to do with "strength" or "being better" (although it can). Fitness is reproductive output -- pure and simple -- and nothing else. It's important to keep these concepts separated.
  • The discussion of speciation can be improved, mainly by introducing the problem of species concepts (and how no species concept is universally useful), and how the most important thing in speciation of sexual organisms is not necessarily geography (allopatry or sympatry), but reproductive isolating barriers of ANY kind. They may be geographic, but they could also be ecological, biochemical, behavioral, etc.
  • The Huxley graphic showing the skeletons of hominids is all right, but it unfortunately resembles all-too-closely the kind of iconic left-to-right linear evolutionary "progress", that doubtlessly causes Steve Gould to roll over in his grave, and will cause me to do so as well when my time comes. The image presented here is not exactly the kind of "linear progress" graphic that is so common, but I am sure we could find a much better graphic to illustrate the concept of *homology* being the signature of evolutionary descent.
  • A lot more can be said in the "History of Evolutionary Thought" section -- specifically, on the kinds of *observations* that had been around for years, that were consistent with Darwin's explanation. For example, the Linnean system of classification predates Darwin and knows nothing of common ancestry and descent, yet its structure as a set of "nested groups" very neatly reflects the true branching nature of the history of life.
Even the main article on this subject looks like it could use some work. There seems to be confusion about what evolutionary ideas predated Darwin and how fast his ideas were accepted. Gnixon 02:38, 28 March 2007 (UTC)Reply
  • Some fleshing-out of the very true statement that "evolution is the organizing principle of all biology" would be justified on this page.
Here's the sort of topic that could really benefit from an expert's perspective. Gnixon 02:38, 28 March 2007 (UTC)Reply

There are some ideas. Take 'em or leave 'em. I'm willing to help, as long as the debate and round-&-round is kept to a minimum. Kind regards, Mandaclair 19:53, 27 March 2007 (UTC)Reply

Excellent suggestions from an expert in the field. Thanks, Mandaclair. Let's get to work! Gnixon 20:20, 27 March 2007 (UTC)Reply
Thanks. (Mandaclair)

Cooks in the pot

More students using Wikipedia as authoritative source. Experts may be discouraged from contributing by "too many cooks in the pot."

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A final note, for now (from User:Mandaclair): The main reason I have taken an interest in this article, is because University students are using Wikipedia more and more as an authoritative source -- a fact that is potentially exciting on one hand, and terrifying on the other. As someone who interacts with biology majors on a daily basis, it would make my job (and my colleagues' jobs) much easier if we helped out in making popular resources (like Wikipedia) as accurate as possible. Otherwise, we spend a lot of time helping students "unlearn" what they thought was true about Evolution (such as: it's all adapation, or it's all a directional process of improvement, or the notion that simply because we refer to "evolutionary theory", that therefore evolution must be some kind of tentative hypothesis that has not been "proven" one way or another... you get the picture.)

Unfortunately, I am sure that many academics in many fields are deterred by the too-many-cooks environment at Wikipedia, and yet, they may feel compelled to help out in some way -- especially if their students use Wikipedia. All of that being said, the Evolution article (as it stands now) does cover most of the main points, and is a decent introduction to the field and its concepts. It could just be a lot clearer, a lot more accurate on some fundamental points, and it could cite more (and better) examples, in many places.

There really are a lot of cooks stirring this pot, but a little word of mouth around the department could go a long way toward increasing the proportion of master chefs. I've wondered sometimes about the idea of creating, for example, a "HarvardBiologyProf" account on Wikipedia that could be shared by a number of experts who each have limited time available. (BTW, I'm not a Harvard bio prof.) Gnixon 04:22, 28 March 2007 (UTC)Reply

Thanks, and for now I think I'll leave most of the editing to the more passionate editors here -- I'm happy to help upon request, Mandaclair 20:54, 27 March 2007 (UTC)Reply

Natural selection

Proposed definition of natural selection. Criticism of adaptationist tone in article. Importance of superfecundity.

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Here are some of the other changes I made last night, and the rationale (Comments by User:Mandaclair.):

  • Natural selection, one of the processes that drives evolution, is a self-evident mechanism that results from the difference in reproductive success between individuals in a population. Natural selection occurs due to two biological facts; 1.) the existence of natural variation within populations and species, and 2.) the fact that all organisms are superfecund (produce more offspring than can possibly survive.) In any generation, successful reproducers necessarily pass their heritable traits to the next generation, while unsuccessful reproducers do not. If these heritable traits increase the evolutionary fitness of an organism, then those organisms will be more likely to survive and reproduce than other organisms in the population. In doing so, they pass more copies of those (heritable) traits on to the next generation, causing those traits to become more common in each generation; the corresponding decrease in fitness for deleterious traits results in their become rarer.[1][2][3]

The important thing about selection is that it is a *self-evident* process, in that: given the undeniable, observable biological facts that 1.) organisms vary, 2.) most variation is heritable 3.) organisms produce more offspring than can possibly survive, and 4.) some heritable traits will influence reproductive success, it *necessarily follows* that heritable traits that increase reproductive success will increase in frequency, while heritable traits that do not increase reproductive success will decrease in frequency or disappear entirely. This is why a very common reaction in the scientific community to the publication of The Origin, was basically along the lines of: "well, DUH, how come *I* never thought of that?" It is self-evident to any thinking, rational human.

Also, it is tempting to think of all evolution and natural selection as "adaptation to the environment", but that is a somewhat naïve point of view, mainly in that it is incomplete (many traits are preserved due to random factors, or evolutionary constraints that prohibit their disappearance, i.e. genetic linkage or developmental constraints. Adaptation need not enter into the preservation of traits over time.) I strongly recommend toning down the adaptationist tone of this article in general. Natural selection is perhaps best understood if reduced to the self-evident mathematical outcome of perpetuation of certain heritable forms due to the simple fact that there are more copies available to reproduce, and they are better at reproducing. Yes, adaptation occurs, but it is not the driving force. Mutation, drift, and selection are the driving forces.

Also, any discussion on drift *must* point out that drift applies to sexually reproducing organsisms, since drift is generally understood as a result of random matings. Thus:

  • In sexually-reproducting organisms, random genetic drift results in heritable traits becoming more or less common simply due to chance and random mating.

Again, with the concept of speciation and divergence, sexual reproduction must be assumed if you're going to invoke "interbreeding". Many organisms (including eukaryotes) are asexual, and so the ability to interbreed cannot define or describe the divergence process. Thus:

  • With enough divergence, two populations can become sufficiently distinct that they may be considered separate species, in particular if the capacity for interbreeding between the two populations is lost.

Great suggestions. Two things I'm a bit uncertain of: one is superfecundity - organisms certainly don't always produce more offspring than can possibly survive, and that's certainly not required for selection to take place. All that's required is that you do better than your neighbor, as in any race. And two is the above misconceptions section. I was never a fan of its inclusion before, and I don't want to see it making a prominent return. It hurts the article. Graft 19:12, 28 March 2007 (UTC)Reply

Hi Graft -- you are right about "doing better than your neighbor", but superfecundity is absolutely, indispensably part of the mechanism of Natural Selection. It is the reason that survival and struggle for existence becomes an issue. Remember Darwin's argument about the elephants: he picked the LEAST fecund animal he could think of, and reasoned that if all elephants ever born survived and reproduced, the earth would be swamped by them. Here:
The elephant is reckoned to be the slowest breeder of all known animals, and I have taken some pains to estimate its probable minimum rate of natural increase: it will be under the mark to assume that it breeds when thirty years old, and goes on breeding until ninety years old, bringing forth three pair of young in this interval; if this be so, at the end of the fifth century there would be alive fifteen million elephants, descended from the first pair. (Darwin. On the Origin of Species. Ist Ed. Ch 3.)
Even though it was shown later that Darwin got the calculations wrong, his point is still true and *fundamental* to natural selection. And this is elephants! Think of the superfecundity of arthropods, marine non-vertebrates, bacteria, fungi, rodents, plants that reproduce by wind-pollination... the fact of Superfecundity is fundamental to life on earth, to Natural Selection, and to Evolution. In "Recapitulation and Conclusion" (Chapter 14) Darwin also calls superfecundity "a ratio of increase so high as to lead to a struggle for life".
Not sure how related this is, but evolutionary biologists have a term called LRS, "Lifetime Reproductive Success", which is an additive function of the probability of surviving to any given age, times the potential number of offspring that could be produced in each unit of time (such as a year), added up over the entire lifetime of the organism. LRS can never reach infinity, because of selection, deleterious mutations, evolutionary trade-offs, etc. It may help to think of superfecundity at the species level rather than at the level of the individual. You can also think of it this way: if organisms were NOT superfecund, and did NOT produce more offspring than could possibly survive, then there would be no struggle for existence. Selection might result in the *increase* of your neighbor who is "doing better" but without superfecundity it won't result in the *extinction* of those that "do worse". (Comments from User:Mandaclair.)
Ah - as to elephants. This is true and good, but populations frequently do explode and grow in size exponentially, and we can still see the influence of selection in this context - that is, allele frequencies can change as a result of differential fitness (or reproductive ability) in an exploding population. So why would we then say that superfecundity is *indispensible* for selection? Graft 20:58, 28 March 2007 (UTC)Reply
Superfecundity is indispensible because 1.) it is a blatantly obvious fact of biology, and the mechanism of Natural Selection is firmly rooted on this and other biological facts (variation, heritability, superfecundity, survivorship) -- and 2.) superfecundity is the primary reason for the "struggle for existence" in the first place. Also, consider gene flow in a world where there is no superfecundity and thus no struggle for existence. If all variants that are born (hatched, germinated, etc.) *could* survive and reproduce, and there is no struggle for existence, it is hard to imagine how allele frequencies are going to change significantly over long periods of time. Sure there are population explosions but eventually, something's got to give, and it "gives" because THERE ARE TOO MANY INDIVIDUALS, MORE THAN CAN POSSIBLY SURVIVE, GIVEN THE AVAILABLE RESOURCES. Selection *means* selection of certain individuals out of a pool of individuals who can't all "make it" because there are too many of them to all "make it". This is what Darwin believed and what he stated explicitly, and should be included on this page, if only for that reason. It is in Darwin's Introduction, and my quick inspection shows (not surprisingly) that his quotation is already included in the Wiki entry about Darwin:
As many more individuals of each species are born than can possibly survive; and as, consequently, there is a frequently recurring struggle for existence, it follows that any being, if it vary however slightly in any manner profitable to itself, under the complex and sometimes varying conditions of life, will have a better chance of surviving, and thus be naturally selected. From the strong principle of inheritance, any selected variety will tend to propagate its new and modified form.
Here is a lovely link I found showing Ernst Mayr's schematic of Natural Selection. Note that Superfecundity is first principle.
www.scepscor.org/outreach/bio2010/workshop-summary-files/supplemental-material/naturalselection.pdf
Mandaclair 23:11, 28 March 2007 (UTC)Reply
This may be how Darwin defined selection, but as far as I've ever seen it defined, technically, it entirely in terms of differential reproductive success, and nothing else. That's all that's encoded in the idea of fitness. So, while I agree that superfecundity exists and is a fact of nature, I don't see how it is *necessarily* related to selection. Anyway, this is getting abstruse and maybe out of the scope of this article in general. Graft 15:47, 29 March 2007 (UTC)Reply
Well, Ernst Mayr was arguably the most important Evolutionary Biologist of the 20th century, and his schematic of selection (as in the link I've given) has superfecundity as a first principle of selection. Also: technically, Darwin never spoke of reproductive fitness in The Origin using that word (fitness), although differential reproductive success is certainly implied. Note however that fitness is *not* a "differential" (relational) concept in itself. Finally (and this shouldn't be news to anyone), "Survival of the Fittest", in Darwin's time (and meaning) was not a statement of fitness as we define it now -- in The Origin, he really meant survival of competition and, that word you hate, struggle.Mandaclair 23:21, 29 March 2007 (UTC)Reply

Common misconceptions

Continued from Recommendations. Suggestion for section on common misconceptions about evolution. Some support. Concern that such a section would devolve into anti-creationist POV, as did a similar section before.

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Hi -- the last thing I'd like to reply to, is Gnixon's request for more details about "common misconceptions" about evolution. Here's the short list -- some of these may *seem* targeted for the creationists, but they're really not. Even atheists sometimes misunderstand the true meaning of the word "theory". I also realize that many of these issues are addressed piecemeal throughout the article as it stands, but a "bold rewrite" attempt might want to consolidate them into a single section. I think that would be extremely valuable. (From User:Mandaclair.)

  • Evolution is [only] a "theory" that remains to be "proved"
  • Survival of the fittest means survival of the best
  • Human evolved "from" apes (or, any extant X evolved "from" any extant Y)
  • Most of an organism's traits are adaptations for some beneficial function
  • Humans/mammals/vertebrates are the "most advanced" organisms -- everything has been "leading up" to us
  • Evolution always optimizes organisms and improves them over time
  • Evolution is usually a slow, gradual, evenly-paced process
  • The historical path that evolution took was obvious and unavoidable, and how things will evolve in the future can be somewhat predicted

Please e-mail me for questions or details. Thanks, Mandaclair 18:18, 28 March 2007 (UTC)Reply

The only reason I suggest the misconceptions section, is because Evolution is probably the most misunderstood science of all, because it is so prominent and conspicuous in the popular, public eye. Thus Evolution is in a unique position of having to deal with public misconception, more than any other science has to. There is a way to compose a section like this that does not appear like it's catering to creationists, but rather, caters to the very real need to educate and adjust what many people erroneously believe the position of Evolutionary Science to be.Mandaclair 19:37, 28 March 2007 (UTC)Reply
Hi Mandaclair,
I take your point, and believe me I am sympathetic, but this is not an advocacy site; I don't really see the justification for including what's undoubtedly aimed at countering a specific cultural trend here, no matter what the views of the editors. I know others feel differently here, but I think that we should be true to WP, here, not our selves. Graft 20:54, 28 March 2007 (UTC)Reply
Hmmm, I'm not sure how education = advocacy... most of the points I raised are not about advocacy at all, as much as they are about misconceptions people have of Evolution as an optimizing, directional, gradual process of increasing complexification where X evolves "into" Y. I don't need to argue this point any further, but it's a fact that most people view Evolution that way (regardless of their personal "advocacies")... and that view of evolution is thoroughly incorrect.Mandaclair 21:19, 28 March 2007 (UTC)Reply
Misconceptions=Misunderstandings. We used to have a 'Misunderstandings' section but it was thought by many to be WP:POV to have such a section, so it was removed by consensus, on 22 February. There is a separate article called Misunderstandings about evolution, which survived a vote for deletion in January, but its future is still unclear. User:Silence has referred to the title Misunderstandings as 'unacademic and unneutral'. EdJohnston 00:35, 29 March 2007 (UTC)Reply
Interesting... Well, at least this information is still posted somewhere. Cheers, Mandaclair 00:47, 29 March 2007 (UTC)Reply
I support implementing the bulleted points made by Mandaclair above. I also think some mentions of the misunderstandings are needed. As a complex and touchy issue, many people have preconceived notions or blatantly wrong information about evolution, which is a big reason why it has encountered so much opposition. --Hojimachongtalk 01:02, 29 March 2007 (UTC)Reply

Changes implemented

Changes to intro by Mandaclair. Support for them from GetAgrippa.

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Heads up -- I'm going to make a few changes, but none should come as a big surprise. Questions? See archive above.Mandaclair 18:00, 29 March 2007 (UTC)Reply

Wow! Drastic improvement to the intro (also finally corrected the definition to include "time" or "successive generations"). I'd just leave out the last paragraph about history for later. The speciation section could really use the same hand as it is sadly lacking. GetAgrippa 19:46, 29 March 2007 (UTC)Reply
Thanks -- will be tackling Speciation next.Mandaclair 19:49, 29 March 2007 (UTC)Reply

Struggle to survive

Debate over "struggle for survival" phrase as too Victorian, Marxist, anthropomorphic. Defended as accurate description, used by Darwin. Resolution via "roundabout verbage."

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Question, Graft: I see by your edit comment that you "hate the word struggle", but I wonder how much bearing your personal hatred of the word has, given the fact that Darwin consistently used the phrase "struggle for existence" throughout The Origin, and this "struggle" is very much viewed as fundamental to Natural Selection. Seems to me that any description of selection ought to be true to Darwin, at least...Mandaclair 23:07, 29 March 2007 (UTC)Reply

Maybe, but the sentence doesn't really add anything other than flavor. Since you've already expressed an aversion to the adaptationist tone of the article, I'd think you'd be in favor of trimming such sentences. I'm actually quite pleased with the fact that this article has, in general, managed to avoid the "struggle to survive" cartoon of evolution in its language. Most of the positive selection that goes on does not take the form of a visible struggle - it is totally invisible to any observation and can only be detected via statistics or genetics. I dislike that language because it leads people to expect competition - lions snarling over meat, etc. This both presents a distorted picture of evolution and results in misconceptions (like "there is no selection going on in humans right now", because we can't see it). Also, I'll point out that this article has taken great pains to move past Darwin in its language and in its treatment of ideas. Origin of Species is, after all, almost 150 years old now. Graft 03:03, 30 March 2007 (UTC)Reply
The undeniable fact that all organisms struggle for survival (think about it for just a moment -- think about all the energy that is required for all the vital processes. It's no cakewalk) -- has nothing to do with adaptation. The adaptationist perspective is not one of "constant competition and struggle", but one of "every trait is an optimized adaptation for the function it currently serves, and evolution is an optimizing process". Also, even though the Origin of Species was only written about 150 years ago... the principles go back at least 3 and a half billion years. ;-)Mandaclair 03:50, 30 March 2007 (UTC)Reply
Another short note: I find it patently odd that Graft is so opposed to the inclusion of principles that have always been integral to the mechanism of natural selection: namely superfecundity and struggle for existence. I am not aware of any academic reference from a working evolutionary author, alive or dead, that purports to give a complete explanation of natural selection without citing superfecundity and struggle. I think the concept that may be slipping through the cracks here is: natural *selection*, like artificial *selection*, means perpetuation of a *select subset* of the individuals from the previous generation. This *selection* occurs because they cannot all survive. There are too many of them (superfecundity), and life is tough (struggle). This is why it is *selection*. Darwin began his argument for natural selection by thinking and talking about artificial selection. Dachschunds are long and squat because only the long and squat individuals were *selected* for breeding in that lineage, despite the existence of plenty of puppies that weren't long and squat enough. Those other puppies did not become part of the Dachschund lineage. Out of all the bizarre Cambrian animals we find in the Burgess Shale, only a few types were *selected for* and became the modern animal phyla. The rest didn't make it. Superfecundity and struggle. They are part of the process and always have been.Mandaclair 04:11, 30 March 2007 (UTC)Reply
My point about Darwin being 150 years old is that our understanding of these issues has certainly evolved since his time. Case in point, selection. You suggest that *selection* means a select subset of individuals from the previous generation are perpetuated, but this is wrong. *Selection* acts on traits, and more properly acts on allele frequencies. It is an allele that is being *selected* for, and the change in frequency of an allele as a result of differential reproductivity is all that is meant by selection. As I've said before (and which I've yet to see a reply to), superfecundity has nothing to do with this idea. Unless I'm missing something, which it's perfectly possible I am.
At this point it seems to me we're talking about very different things - you're talking about species selection and I'm talking about selection within a species. How to resolve this, I'm not sure, other than to outline both of these ideas.Graft 04:51, 30 March 2007 (UTC)Reply
Selection acts on whole organisms -- not genes, not traits. Yes, traits and genes can be selected FOR or AGAINST, and yes it's all about changing allele frequencies over time, but all of this happens only through the vehicle of the organisms that live, reproduce (or not), and die. There is no other way. Even the "selfish gene" needs the organism to be the phenotypic vessel exposed to natural selection. I will agree that citing the Cambrian was a poor example for me to give, since that is more about interspecific competition, but it was the first thing that came to mind. The principles are clearly applied to the "within a species" level, but I really can't spend any more time trying to justify the rock-solid-established fact that superfecundity and struggle for existence are integral elements of selection, both within a species and for life on Earth in general. The artifical selection example I gave for Dachshunds is perfectly analogous to selection within a species. I am sorry if you don't "see" this point, but you don't have to go 150 years back to Darwin to learn about it. Try looking to Mayr -- he only died a few years ago. I often rode the elevator with him in the Museum of Comparative Zoology... You should have seen him wearing pipe-cleaner ant antennae in the audience on the day of Ed Wilson's last lecture before he went Emeritus...Mandaclair 05:06, 30 March 2007 (UTC)Reply
Also, there's no need to address me in the third person - I can follow along just fine. Graft 04:51, 30 March 2007 (UTC)Reply
Sorry, didn't mean to offend. But since this is the public talk page and not your user page, I figured other folks would be involved in this discussion. My apologies,Mandaclair 05:06, 30 March 2007 (UTC)Reply

I agree with Graft on the issue of "struggle for survival" - it's a metaphor from an earlier age, and it's about as dated as "nature red in tooth and claw". No one talks about species interactions in those terms any more. Guettarda 05:33, 30 March 2007 (UTC)Reply

The funny thing is, I am modern enough to agree with this point of view as well, but only to a certain point. I will be the last person hopping up to paint a picture of "nature red in tooth and claw" and I certainly do not think that "struggle for survival" needs to be *emphasized* greatly when talking about evolution. But it is an *inseparable part* of selection, and of evolution -- not a mere metaphor. That is all I'm saying -- that I can't see justification for leaving it out, but I am 100% on the same page with you that evolution shouldn't be emphasized as some vicious competitive battle out there... although frankly, it really is. This may be hard for humans in industrialized nations to perceive, but do not doubt for a minute that competition for resources among humans worldwide is deadly and fierce. Do not doubt for a minute that organisms by the millions die in floods, droughts, and frosts, that they are consumed by herbivores and predators, that they are driven from their habitats by invasive species, and that they starve to death when a more efficient predator or forager comes along. This shouldn't be emphasized as the central theme of evolution, but it's sheer insanity to deny that it's true.Mandaclair 05:43, 30 March 2007 (UTC)Reply
I'm not saying that the effect of inter- and intra-specific species interactions should be left out, just that calling it a "struggle for survival" is too Victorian, too Marxist, too anthropomorphic a presentation. It also points people in the wrong direction - to think about drastic and dramatic floods, rather than far more mundane features like being shaded out by another individual or killed by a pathogen. Big events don't structure populations nearly as much as do a whole lot of small ones. Guettarda 06:14, 30 March 2007 (UTC)Reply
I dunno, I think pathogens are pretty dramatic too... I may have listed some big dramatic struggles above to make a point, but there are large struggles and small struggles -- even being "shaded out" by another... the main point being that life is never a walk in the park, and there is no free lunch. That's all, and I certainly do agree that we should steer clear of anthropomorphism...but what is another way to word this central concept, other than using the traditional wording "struggle for survival"? Like I said, it doesn't need to be emphasized (at all, and I have never argued for emphasizing it), but it is a key element that I just can't see any reason to justify its exclusion. Any alternative wordings you want to suggest?Mandaclair 06:38, 30 March 2007 (UTC)Reply

It is an old metaphor and the naive also equate it with the survival of the fittest metaphor, but the point I think Mandaclair is making is that biotic competition is a fact of life and superfecundity relates as organisms tend to reproduce more than can survive in any given ecological setting. The terminology maybe a contention but the point does need to be made. I think we would be remiss not to mention both as this is an encyclopedia and the audience needs the basics. Introductory text and books (Gould, Mayr, etc)all mention it to my recollection.GetAgrippa 14:33, 30 March 2007 (UTC)Reply

Yes, I am only arguing for its inclusion for the sake of accuracy and completeness. It is not our fault if readers want to misconstrue this as an "only the swift and the strong shall survive" statement. But just because the concept of a "struggle for existence" may be out of fashion, does not make it untrue.Mandaclair 16:51, 30 March 2007 (UTC)Reply
Sorry to keep harping on this, but I simply don't find this apt in many instances. For instance, let's take skin color. There's strong selective pressures to maintain the right amount of melanin, but they're entirely about reproductive success and nothing else - there's no competition for resources involved, and there's no struggle against other members of the same or any other species. I don't think struggle is an appropriate metaphor for evolution *in general*, and the language above doesn't make me any more inclined to believe it's a useful way of phrasing things. Graft 18:33, 30 March 2007 (UTC)Reply
Graft, "struggle for existence", or whatever you'd like to term the fact that "life is not easy and requires a great investment of energy", is a first-principle because of superfecundity: more offspring are produced than can survive in a world with limited resources. It is as simple as that, the concept has always been central to Selection and Evolution (and Ecology: please recall K, carrying capacity) -- and I personally am tiring of this argument. Achieving any kind of reproductive success always implies struggle, in terms of energy expenditure, acquiring resources, access to mates (in sexual organisms), and biological investment in reproduction. Whether the trait you're looking at is skin color or anything else: if selection favored it, it necessarily implies that individuals carrying the trait were selected FOR and those that didn't carry it were selected AGAINST, and not because life is a bowl of cherries available for the taking. Whether or not you like the word struggle or the concept of struggle: maximizing your fitness IS AN UPHILL BATTLE, and individuals that are better at it persist, while others will not. It doesn't require invoking hand-to-hand combat, tribal wars, and "quarreling over the kill" as being connected to every single trait. Can we please table this topic until we hear a few more views, and until someone bothers to review the primary modern literature that describes Natural Selection. And may I please remind you: nobody is suggesting including anything in this article about "struggle", other than mentioning it briefly as one of the first principles of natural selection. Thanks Mandaclair 19:35, 30 March 2007 (UTC)Reply
Proposed solution:

In an attempt to dress up an old concept in less Victorian/anthropomorphic language, I have gone ahead and replaced the classic "struggle for existence" phrase with some roundabout verbage that, to my mind, means exactly the same thing: "organisms in a population are not all equally successful in terms of survivorship and reproductive success". Conceptually, it is identical to "struggle for existence" -- does this wording satisfy the dissenters?Mandaclair 19:58, 30 March 2007 (UTC)Reply

Good edit, Graft -- I dig, Mandaclair 20:22, 30 March 2007 (UTC)Reply

Variation and Heredity

Call to cut Variation and Heredity sections. Some support for only summarizing variation and heredity within another section. Is adaptationist perspective a POV issue? How is evolution taught these days? Few comments.

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I kind of feel like the short "Variation" and "Heredity" sections don't belong here (mainly because the way they are written does not really address Evolution). What do folks think about deleting these sections -- keeping in mind that there will be embedded links to the variation and heredity articles, throughout this one?Mandaclair 04:24, 30 March 2007 (UTC)Reply

I would agree with a rewrite of Variation and Heredity, but variation (mutations and recombinations)and heritable need to be explained just like superfecundity should be mentioned. It doesn't have to have a separate section. I tend to agree with your analysis of adaptation, but that is a POV issue (I can see the Gouldian influence in your education)as many authors emphasize adaptation. I do agree that exaptations should be mentioned. I am curious how evolution is taught nowadays (it has been over twenty years since I taught an introductory biology course and molecular biology and genomics has drastically altered the state of affairs).GetAgrippa 14:33, 30 March 2007 (UTC)Reply
Here's how evolution is (mostly) taught today: phylogeny, phylogeny, phylogeny! Students get the fundamentals and the history of the field... but then a lot about evolutionary genomics, evo-devo, gene & genome duplications, etc. As you might expect...Mandaclair 17:05, 30 March 2007 (UTC)Reply

Selection and Adaptation

Edits by Mandaclair to Selection and Adaptation section. Brief debate over ecological selection.

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I have now made some bold(ish) edits to the Selection and Adaptation section, a bit more consistent with the way these concepts are taught in Evolution courses for biology majors. The previous version of this section was really a bit off... for example, the 3rd mode of selection is disruptive selection (not artificial selection), and all 3 modes could be argued to select against harmful traits and select for beneficial ones. I also tried to improve the description of sexual selection a bit, and removed the distinction of "ecological selection" because it seemed a bit redundant with the existing description of natural selection in general. "Ecological selection" is not a term I hear used a lot... it makes sense, sure, but I don't think it's any kind of standard category of selection... As I go through this article, though, I am generally very impressed with its quality. My intention here is just to tidy-up, not do any drastic rewrites! Thanks, Mandaclair 05:20, 30 March 2007 (UTC)Reply

I also have to disagree with you on ecological selection. I'd say it has a lot of use in the last 5 years. I'd say it's at the very least presented as a distinct type of selection - e.g., [2], p.127. Guettarda 06:16, 30 March 2007 (UTC)Reply
I know Patrick, and in fact worked with people in the Shaw lab for many years. In that paper, I think the term "ecological selection" is used mainly as a convenience (in context) to distinguish it from sexual selection in the argument they are making. That's just my opinion, but I do have to say that, although the term "ecological selection" certainly makes sense, I don't hear it used often as *its own term* (most people just say natural selection and sexual selection, or talk about the 3 modes). I do note, as you say, a lot more recent usage of this term... My only objection was the prior categorization scheme in the article, which divided selection first into "ecological" and "sexual", and then later into "directional", "stabilizing", and "artificial"... the divisions were somewhat confusing. But if more people here think ecological selection belongs in the article in some way, I say put it back, as long as it is implemented in a way different from that previous categorization. In my opinion "ecological selection" is not in such common use that it warrants status as a category of selection in this article... (e.g. ecological selection, in quotes, gets about 20,000 google hits, while sexual selection in quotes gets over 900,000... not really terms or categories in equal usage) Thanks, Mandaclair 06:38, 30 March 2007 (UTC)Reply


A note on the system of boxes just above (used for Mandaclair's comments and the responses): User:Gnixon is the one who wrote the summaries and created the system of boxes. (It would be more clear if he would add his own signed comment to announce the refactoring). In fact, it does save space on the Talk page, and I like the system, but perhaps not everyone does. Please respond here either for or against this type of refactoring. I think there is a consensus that it should be done for questions answered in the FAQ, but there is not yet a consensus for doing it more generally. There is a sub-question as to whether some further action should be taken on Mandaclair's suggestions. Respond here on that issue as well, if you have an opinion. EdJohnston 16:53, 3 April 2007 (UTC)Reply

For' (obviously). You're right, Ed. I should have said something about it. I certainly hope the archiving and subject headings haven't stifled discussion, but it was getting so long and covering so many topics that I couldn't follow things anymore. Gnixon 20:55, 3 April 2007 (UTC)Reply
For so long as its employed sparingly to avoid confusing new/casual readers. But it is so obviously useful for high traffic talk pages such as this; I hope to utilize it elsewhere. - RoyBoy 800 23:14, 9 April 2007 (UTC)Reply

Lead

Recent changes to the lead seem to have been well-received, but I think they've also exacerbated an existing problem: the lead is far too long and detailed.

WP:LEAD recommends that the lead be concise and accessible, and suggests that it should be between one and four paragraphs long. The current lead is 7 paragraphs long, and I think one could easily argue that its neither concise nor accessible to the average reader. What's more, from glancing at the table of contents, the lead hardly seems to be an "overview" of the article. (Granted, the article's contents are not well organized.) Some articles about major scientific fields have addressed the issue by including only the definition in the lead, then following with an "Introduction" section. I'm not sure that's the best solution, but we have to do something. Any ideas? I'll try to make a content-neutral revision sometime soon unless someone beats me to it. Gnixon 18:39, 31 March 2007 (UTC)Reply

A survey about the lead took place here. Thanks, Ed, for mentioning it. Gnixon 18:58, 3 April 2007 (UTC)Reply

Reviewers during January's FAR stressed that this article needed work on being accessible to its readers, especially in the intro. See FAR section below. Gnixon 19:39, 3 April 2007 (UTC)Reply

Organization

A glance at the table of contents is enough to prove that this article has become very poorly organized. I'd like to undertake a major reorganization, one that is content-neutral but better sorts things under headings and subheadings. I think a similar change at Physics worked out well (compare before [3] and after [4]). I'd appreciate some input regarding what the table of contents should look like and what goes where. Thanks! Gnixon 18:39, 31 March 2007 (UTC)Reply

I would need to review User:Silence's previous plan, and the feedback questionnaire that he created for the lead, to get some ideas. (It's all in this Talk page or the archives). He also made a list of issues he thought would need to be fixed to get back the FA status. I can try to dig up all the diffs pointing to that stuff later. EdJohnston 16:57, 3 April 2007 (UTC)Reply
The lead survey is here. Gnixon 18:53, 3 April 2007 (UTC)Reply
The featured article review, including Silence's extensive comments, is here. Gnixon 18:56, 3 April 2007 (UTC)Reply

FAR

The featured article review in January resulted in delisting, but also produced a number of well-received recommendations from User:Silence and others. Not all of them have been carried out. I've copied Silence's list of recommendations in the hidden archive below. Please comment either within the archive or below it. Gnixon 19:28, 3 April 2007 (UTC) Reply

Silence's FAR recommendations.
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25 problems to resolve, for starters:

  • The third paragraph of the lead section goes into too much detail about the circumstances surrounding The Origin of Species. That level of detail might be appropriate in the "History" section, but no more than a few words should be devoted to it in the lead section.
  • The armadillo image has an excessively long caption, bloated by trivia. It is also poorly-placed; having two lengthy vertical blocks of text and image at the top of the article makes the page look clumsy and cluttered. The armadillo thing should probably be either shortened and transplanted to another part of the article, or removed altogether.
  • Section titles should not be capitalized. "Basic Processes" and "Mechanisms of Evolution" are thus incorrect.
  • It is incorrect to italicize "e.g." and "i.e.". (There is also some excessive and inconsistent use of the latter.)
  • It is incorrect to italicize quoted text.
  • Some languages crosses the line from being simple and user-friendly to being overcasual. Academic, encyclopedic tone should be maintained, and we should avoid treating our readers like infants with phrases like "phenotypic variation (e.g., what makes you appear different from your neighbor)".
  • Although the article does a good job of explaining most terms, some new terms are still unexplained, and a surprising number are unlinked, like gene, genotype, genetic variation, and many more.
  • There is an overuse of parentheses in this article. These can be replaced by em-dashes, commas, etc. in some cases, to avoid making the text seem fragmented.

*Avoid external links in the article text, like the Tetrahymena link.

  • There are various minor grammatical errors that are not significant enough to mention here; a thorough copyedit should fix them.
  • "Selection and adaptation" seems to be a little too long and a little too listy, relative to the other, more compact sections. Cutting down on all the subtypes listed could probably cut this section's length almost in half; that level of detail is more appropriate for the daughter articles anyway. This section also needs references, badly—especially for its paragraph on evolutionary teleology.

*Bolding should not be use to emphasize a random word in a prose paragraph.

*There is some inconsistency in reference style in sections like "Cooperation".

  • There is poor illustrative balance in the "Evidence of evolution" section. All three images deal with aquatic animals, suggesting to uninformed readers that there isn't any evidence for evolution from other species; this impression should obviously not be implied, so at least one of the images should be removed, and other images should be added. The "nasal drift" image seems like the least useful one at the moment; although it's very pretty, the sequencing and similarity is least obvious.
  • Considering how drastically the rest of this article has been shortened, you may want to consider shortening the "Evidence of evolution" section too, to avoid imbalance. This can be easily done by cutting down on examples and trivial details.
We should also consider what the point is in having "Evidence of evolution." Is there a better title, like maybe "Examples of evolution"? Gnixon 19:53, 3 April 2007 (UTC)Reply
  • "History of the modern synthesis of evolutionary thought" should clearly be a subsection of "Study of evolution", and should be shortened to a simpler title, like "History of evolutionary thought".
  • The "History" section is currently far too short. Important information that was removed should be re-added to make it at least 50% bigger ("Academic disciplines", below, is a good example of a nice-sized section). To give an idea of how much compression is appropriate, 3-5 fair-sized paragraphs (about 4 sentences in length each) should be the goal. Anything much shorter or longer than that is not appropriate.
We're still on the short side of things here. There are only about 2 paragraphs on history. The section appears long because it has a subsection "Academic disciplines". Surely that would work better as "Current research". Gnixon 19:53, 3 April 2007 (UTC)Reply
  • The "Misunderstandings" section is too short, and some very important information (e.g. about the fact-theory distinction) has recently been removed from the article, making it much less informationally valuable to readers. Of course, whether a "Misunderstandings" section (or its new daughter article) is appropriate here at all should be discussed; there is little precedent for such a move, and it seems to fly in the face of academic and NPOV conventions, as well as to be a very useless categorizational method--a misunderstanding about the nature of mutations, for example, would be very useful if put under "Mutations", but useless if put under the generic heading of "Misunderstandings". Ideally, thus, a "Misunderstandings" section should simply be split up into sections dealing with the specific topics involved in each misunderstanding. From an NPOV perspective, it is particularly troubling to see statements to the effect that the creationist movement was caused by misunderstandings of evolution; it is perfectly fair to say that creationists regularly misunderstand evolution, but to make inferences and judgments from that is not NPOV; at the very least, such statements should be replaced with attributed ones, so it is not Wikipedia itself that is making them.
Misunderstandings was cut. It spawned a Misunderstandings subarticle and the "Objections to Evolution" subarticle, which are mentioned in the Social controversy section. Others have independently argued that a Misunderstandings section would be useful, but still others have expressed concerns that it's too vulnerable to POV. I wonder if we could reintroduce "Misunderstandings," but excercising *extreme* care to avoid anything about creationism? Gnixon 19:53, 3 April 2007 (UTC)Reply
  • This article needs to have a "social effects" section. The effects of the study of evolutionary biology on society and culture over the last few hundred years is immense, and highly noteworthy. This would be a more appropriate and useful place to (briefly) discuss creationism than a POVed "misunderstandings" section, obviously.
Implemented. This might make an NPOV "Misunderstandings" section more feasible. Gnixon 19:53, 3 April 2007 (UTC)Reply
  • The "See Also" section is too large. Ideally, there should be no "See Also" section at all for a time-level article like this; any highly important articles should be mentioned in the article text and/or series templates, and any less important ones should not be mentioned in this article, but rather in daughter articles. Some of the articles listed here are not even real articles, like Animal evolution.

*Why is there an empty "Notes" section?

  • A number of the references are broken or inconsistent. It will take an in-depth review and copyedit to make them all consistent.
  • The external links should be cut down a little. 10-15 is ideal for an FA; there are currently 20. One good method to shorten the list without removing important information is to simply use some of the links in the "References" section; this gives them the added value of having relevance to specific parts of the article, as opposed to just being "add-ons". EL section reduced to 9 items in February. EdJohnston 20:04, 3 April 2007 (UTC)Reply
  • Is there any particular reason that Evolution, rather than Modern evolutionary synthesis, is under Category:Theories?
-Silence 19:58, 6 January 2007 (UTC)Reply

The references have been corrected, broken links removed, internal links were formatted according to WP:CITET, and outside references have been shortened. Other activities to make this an FA are required. Orangemarlin 19:31, 3 April 2007 (UTC)Reply

For technical items where it's absolutely clear they've been resolved, I suggest striking through the items. Gnixon 19:53, 3 April 2007 (UTC)Reply
Gnixon, thanks for digging up this useful info. I struck out Silence's action item about external links, since they were reduced to nine back on 9 February. If I see more things I can fix I'll edit the boxed copy of his list you provided above. EdJohnston 20:09, 3 April 2007 (UTC)Reply
I also struck out some items that were completed. —The preceding unsigned comment was added by Orangemarlin (talkcontribs) 20:12, 3 April 2007 (UTC).Reply
AOL, I'd like to strike out the infant issue. I agree with the idea of not over explaining concepts in article, but I would emphasize that a "phenotype" is not common knowledge. - RoyBoy 800 23:42, 9 April 2007 (UTC)Reply
Fine to strike it out if it's no longer a problem, but the complaint was justified: "e.g., what makes you different from your neighbor" is far too casual language for an encyclopedia. Gnixon 19:19, 10 April 2007 (UTC)Reply

Language

After making extensive changes to help resolve items on his list, as well as making improvements to several related articles, Silence left this comment:

Unfortunately, I lack the biological expertise to fix some of this article's largest problems: the opaqueness of some of the more technical sections, lacking even an attempt to provide readers with context in many cases, rnders large portions of this article essentially useless as a general reference tool. What we need is some more work on clarifying concepts by people who are both very familiar with the processes and mechanisms involved, and able to explain them in sufficiently clear, engaging language. We need a Dawkins! :( -Silence 06:25, 6 February 2007 (UTC)

He also voted "remove" for similar reasons:

Remove unless dramatic improvements ensue. I can only do so much; the incredibly confusing mess of various parts of the "processes" and "mechanisms" sections will require a substantial rewrite by knowledgeable folk in order to be of any use to readers; there's nothing wrong with using complex concepts and important technical terms, but the article's frequent failure to keep its readership in mind and coherently explain these things, as well as poor writing quality in a number of paragraphs and inconsistency in references, makes the current article unfit to be an FA. Hopefully, if efforts aren't rallied beforehand, they will become more focused as a result of the demanding pressures of the FAC and peer-review process. -Silence 06:25, 6 February 2007 (UTC)

Other reviewers also stressed the need to explain concepts in accessible language, especially in the Intro. Gnixon 19:37, 3 April 2007 (UTC)Reply

You and I, given that we have strong disagreement on many issues pertaining to these articles, cannot be the only two who are involved. I would "hold your horses" until other editors weigh in with their opinions. You have a tendency to go "ready, fire, fire, fire, aim." Slow down. Orangemarlin 19:58, 3 April 2007 (UTC)Reply
Whereas your tendency is "revert, revert, then maybe read."  ;-) Just trying to be bold until there seem to be objections. I haven't yet changed anything about the article. I think the area where we disagree is pretty well-defined, so we can probably cooperate on other things. It's a shame that there haven't been many editors around here lately. Gnixon 20:27, 3 April 2007 (UTC)Reply
By the way, what happened to the lead? It is way too long. I think it grew by creeping. Orangemarlin 19:59, 3 April 2007 (UTC)Reply
Yes, indeed. See my comments above. Gnixon 20:27, 3 April 2007 (UTC)Reply

I have said over and over that this article needs to be accessible. Unfortunately, that seems to be a very difficult thing to achieve.--Filll 20:04, 3 April 2007 (UTC)Reply

We're trying. Come back and help, this article needs you too. Orangemarlin 20:11, 3 April 2007 (UTC)Reply

Semantics

I'm not seriously proposing a move, but can anyone see how many of the problems we have here would be solved by changing the article name to "Evolutionary biology"? Many of the tensions on this page are due to confusion over whether we're writing about

  1. Evolutionary biology, a field of study like Physics.
  2. Theory of Evolution, as in, the Modern Synthesis, a theory like the Theory of General Relativity
  3. Evolutionary processes, as in the observable aspects
  4. Evolution by natural selection, meaning the concept of it, as in Darwin's revolutionary idea that changed science and society, like Adam Smith's Invisible Hand.

How did the English language come up so short here? How do other encyclopedias handle the problem? Gnixon 20:49, 3 April 2007 (UTC)Reply

Let's keep the name Evolution. The idea of changing the name comes up promptly every six weeks, and is always rejected. The name has been this way since 2001. EdJohnston 14:05, 5 April 2007 (UTC)Reply
Hey, I said I wasn't proposing a move!  :-) On the other hand, I think it's worth discussing which definition we're writing about, or which parts of the article address each meaning. I also think we have almost enough material to make a separate "Theory of Evolution" article, and I wish we had enough to make "Evolutionary biology" (as in the branch distinguished from molecular bio). Gnixon 14:50, 5 April 2007 (UTC)Reply

LUCA again

FYI, apropos our debate of a week or two ago, I today read a bit by Doolittle (and Eric Bapteste) about the Tree of Life, in PNAS, Feb 13 2007, titled "Pattern Pluralism and the Tree of Life hypothesis". He uses some strong language which I have no doubt will end up in some creationist quote mine (cf. his first sentence, "The meaning, role in biology, and support in evidence of the universal ‘‘Tree of Life’’ (TOL) are currently in dispute." Good read. Graft 22:37, 3 April 2007 (UTC)Reply


Excellent, thanks Graft. I remember at one point in the past evolutionary webs was emphasized more than trees, but apparently it is a little of both. I agree this will end up in creationist quote mines. Doolittle admits that tree patterns suffice for most of life and that he is referring predominately to prokaryotes because of HGT and fusion events. He gets rather philosophical also (which I tend to agree with some of his sentiments but don't agree with reaction of stifling the whole pursuit). I do think that Doolittle is fatalistic about it as others disagree:

Kurland CG, Canback B, Berg OG. Horizontal gene transfer: a critical view. Proc Natl Acad Sci U S A. 2003 Aug 19;100(17):9658-62. Epub 2003 Aug 5. Review. PMID: 12902542 [PubMed - indexed for MEDLINE]


Lake JA, Rivera MC. Deriving the genomic tree of life in the presence of horizontal gene transfer: conditioned reconstruction. Mol Biol Evol. 2004 Apr;21(4):681-90. Epub 2004 Jan 22. PMID: 14739244 [PubMed - indexed for MEDLINE]


Ge F, Wang LS, Kim J. The cobweb of life revealed by genome-scale estimates of horizontal gene transfer. PLoS Biol. 2005 Oct;3(10):e316. Epub 2005 Aug 30. PMID: 16122348 [PubMed - indexed for MEDLINE]

Kurland CG. What tangled web: barriers to rampant horizontal gene transfer. Bioessays. 2005 Jul;27(7):741-7. PMID: 15954096 [PubMed - indexed for MEDLINE

I think Doolittle is correct to throw a red flag of reasonable doubt, but it should be a cautionary tale to proceed with caution rather than render it mute. In the end, the article will be used by creationist quote mines that another Darwinist scientist disproves evolution theory is feasible. GetAgrippa 14:08, 4 April 2007 (UTC)Reply

Sounds like stuff that would fit very nicely into a "current research" section. (Also, I wouldn't sweat the creationist angle too much. It's always easy to distinguish between arguing the details and arguing the big picture.) Gnixon 14:22, 4 April 2007 (UTC)Reply

Doolittle's last paragraph deals with the issue of Creationists:
Holding onto this ladder of pattern is an unnecessary hindrance in the understanding of process (which is prior to pattern) both ontologically and in our more down-to-earth conceptualization of how evolution has occurred. And it should not be an essential element in our struggle against those who doubt the validity of evolutionary theory, who can take comfort from this challenge to the TOL only by a willful misunderstanding of its import. The patterns of similarity and difference seen among living things are historical in origin, the product of evolutionary mechanisms that, although various and complex, are not beyond comprehension and can sometimes be reconstructed.
But I do think his point should be well-taken, that one shouldn't assume a rooted, branching tree extending back to the beginning of life when we have no way of showing that this must be the case for the deepest parts of the Tree of Life. I haven't read the above HGT review yet, but what do you think of Doolittle's central point - that the assumption that there's something to be identified beyond all that HGT is unfounded to begin with? Graft 16:55, 4 April 2007 (UTC)Reply

Adam has made some good edits regarding LUCA and HGT. Gnixon 12:38, 7 April 2007 (UTC)Reply

Suggested Edit

As a biologist, I have issue with the use of word 'design' in the following sentence in the 'Academic Disciplines' section: The capability of evolution through selection to produce designs optimized for a particular environment has greatly interested mathematicians, scientists and engineers. Could 'design' be replaced with 'biological processes and networks' or something similar? Evolution doesn't generate function through 'design' but with whatever paradigm works.

Thanks! —The preceding unsigned comment was added by 69.181.191.134 (talk) 06:18, 5 April 2007 (UTC).Reply

Done. Thanks for the notice. By the way, you can create an account and edit this article.--ĶĩřβȳŤįɱéØ 06:24, 5 April 2007 (UTC)Reply

Fisherian runaway

I've done a cleanup on Fisherian runaway and trimmed it somewhat. Can someone do a sanity check and make sure I haven't removed anything important? Also, it would be good if someone could add some references to it. (I'm posting here because Fisherian runaway is pretty low traffic.) Regards, Ben Aveling 08:27, 7 April 2007 (UTC)Reply

BNME:Monkeys

bnme:this is myvirgn attmpt at a internet talk site. I can't type and I have to soetimes hit the keys twice to get em to work. If I a using someones post, please let me k(twice)now. With these thumbs I save monkeys. There had to be some miracle.—The preceding unsigned comment was added by Bnmeee (talkcontribs) 08:52, 7 April 2007 (UTC).Reply

The lead

...My god, what happened to it? It's not a summary of evolution any more, it's back to using undefined jargon (genetic drift is *NOT* a term you can just drop into the lead without comment, and is generally completely inappropriate.

I tried to raise the issue a few topics up, and there was no response. We seem to have a lot of editors here who are interested in the details of the theory, but not many who are interested in good writing. By the way, I explained there "what happened to it." Gnixon 12:53, 7 April 2007 (UTC)Reply

...Really, I don't see how this article is ever going to reach FA again at this rate. For every step forwards, someone turns around and makes in incomprehensible to non-biology majors again. Does anyone really expect a layperson to understand the second paragraph with talk of the Hadean era, RNA world, and so on? Adam Cuerden talk 11:57, 7 April 2007 (UTC)Reply

Here's the old verson of the lead. Maybe this and the new version can be combined into something useful? I dunno. I'm tired of this nonsense. It seems like every month a new simple lead gets made, then someone replaces it with an incomprehensible one.

In biology, evolution is the process of change with time in the inherited characteristics, or traits, of a population of organisms. Heritable traits are encoded in the genetic material of an organism (usually DNA). Evolution results from changes in this genetic material (mutation) and the subsequent spread of these changes in the population, and explains the observed changes over time in the fossil record.

Natural selection, one of the processes that drives evolution, results from improved reproductive success by individuals best adapted to survive and reproduce in a given environment. These successful survivors and reproducers pass their beneficial, heritable traits to the next generation. If these traits increase the evolutionary fitness of an organism, they will be more likely to survive and reproduce than other organisms in the population. In doing so, they pass more copies of those (heritable) traits on to the next generation, causing advantageous traits to become more common in each generation; the corresponding decrease in fitness for deleterious traits results in their becoming rarer.[1][2][3] In time, this results in adaptation: the gradual accumulation of new beneficial traits and the preservation of existing ones results in a population of organisms becoming better suited to its environment and ecological niche.[4]

Though natural selection is decidedly non-random in its manner of action, other more capricious forces have a strong hand in the process of evolution. Genetic drift results in heritable traits becoming more or less common simply due to random chance. This aimless process may overwhelm the effects of natural selection in certain situations (especially in small populations).

Differences in environment, and the element of chance in what mutations happen to arise and which ones survive, can cause different populations (or parts of populations) to develop in divergent directions. With enough divergence, two populations can become sufficiently distinct that they may be considered separate species, in particular if the capacity for interbreeding between the two populations is lost. Evidence such as the wide distribution of the canonical genetic code indicates that all known cellular organisms are ultimately descended from a common ancestral population.[1][5][6][7][8][9][10][11]

While the idea of evolution (as opposed to the fixity of species) is ancient, the modern concept of evolution by natural selection was first set out by Alfred Wallace and Charles Darwin in a joint paper to the Linnean Society, followed by the publication of Darwin's 1859 book, On the Origin of Species. In the 1930s, the modern evolutionary synthesis combined Darwin's natural selection with Gregor Mendel's genetics. As more and more evidence was collected and understanding of the processes of evolution improved, evolution became the central organising principle of biology.[12][13]

Even the old version is too long by WP:LEAD standards. I really think a big problem is that editors can't decide if we're writing about evolution in general or about the details of the theory. I don't think natural selection and speciation need to be explained in nearly so much detail, and I don't think genetic drift needs to be mentioned at all (in the intro), but clearly other editors disagree. I'd love to see an expert (I don't qualify) try to write a concise 2-4 paragraph lead (not even "introduction", just "lead") that covers the big ideas in a readable, engaging way. Gnixon 12:53, 7 April 2007 (UTC)Reply

I always get a little defensive when people complain about Lead length. Lead length should never be an issue so long as the lead does indeed accurately summarize an article. If the article is long, detailed and conveys a complicated subject... I'm sorry but the lead had damn well better be long! :"D I get defensive, because a comprehensive (rather than light introductory) lead is essential; since, as I understand it, some projects propose to just use the leads of articles for various publications/replications. - RoyBoy 800 00:01, 10 April 2007 (UTC)Reply
That's an interesting point. It often seems to me that big articles would do well to have something even more concise than WP:LEAD, followed by some sort of "Introduction" before getting to other topics. Maybe a good analogy would be the dustjacket and introduction of a book. The dustjacket has to be brief enough to grab the reader's attention, but an introduction has a beginning, a middle, and an end, and can be quite long if necessary. Gnixon 00:09, 10 April 2007 (UTC)Reply
Sounds about right. Although not necessarily relevant, I have been heavily involved in shaping two leads. Abortion because it was mired in debate, and Blade Runner because I wanted it featured (after I had substantially increased the article size). I think the Blade Runner lead is a solid example of what a lead should be for beefy articles; and I actually had a minor disagreement with a user who was under the impression leads should be a few sentences long. If I remember correctly that helped tighten up the lead enough to then create the 4th paragraph. Um.... I should stop rambling now. - RoyBoy 800 00:26, 10 April 2007 (UTC)Reply
Pointedly abortion is very short, and that does make me consider expanding it to summarize the great article underneath... but I'd need to ditch my real life to make that happen. :"D RoyBoy 800 00:31, 10 April 2007 (UTC)Reply
Something we did for abortion is to create a subpage for debate and discussion of the lead, which helped focus things a great deal. Then when everything had been said and differences in opinion had solidified; many versions were created, criticized and rewritten, again and again in subsections to provide a chronology and direction for what the lead eventually became. - RoyBoy 800 00:43, 10 April 2007 (UTC)Reply
As someone who used to edit here regularly, I must say the lead has gone seriously downhill. Step one: just put the old lead (from ~mid 2006) back and then try to make it a bit more user friendly. The current lead is just horrible. Mikker (...) 13:09, 7 April 2007 (UTC)Reply


Here's the mid 2006 lead. It has one major problem in paragraph 2 - too many unexplained terms - but I think something could be made of it:

In biology, evolution is the change in the heritable traits of a population over successive generations, as determined by shifts in the allele frequencies of genes. Through the course of time, this process results in the origin of new species from existing ones (speciation). All contemporary organisms are related to each other through common descent, the products of cumulative evolutionary changes over billions of years. Evolution is the source of the vast diversity of extant and extinct life on Earth.[14][15]

The basic mechanisms that produce evolutionary change are natural selection (which includes ecological, sexual, and kin selection) and genetic drift; these two mechanisms act on the genetic variation created by mutation, genetic recombination and gene flow. Natural selection is the process by which individual organisms with favorable traits are more likely to survive and reproduce. If those traits are heritable, they are passed to succeeding generations, with the result that beneficial heritable traits become more common in the next generation.[16][17][18] Given enough time, this passive process can result in varied adaptations to changing environmental conditions.[4]

The modern understanding of evolution is based on the theory of natural selection, which was first set out in a joint 1858 paper by Charles Darwin and Alfred Russel Wallace and popularized in Darwin's 1859 book The Origin of Species. In the 1930s, Darwinian natural selection was combined with the theory of Mendelian heredity to form the modern evolutionary synthesis, also known as "Neo-Darwinism". The modern synthesis describes evolution as a change in the allele frequency within a population from one generation to the next.[4]

The theory of evolution has become the central organizing principle of modern biology, relating directly to topics such as the origin of antibiotic resistance in bacteria, eusociality in insects, and the staggering biodiversity of the living world. The modern evolutionary synthesis is broadly received as scientific consensus and has replaced earlier explanations for the origin of species, including Lamarckism, and is currently the most powerful theory explaining biology.

Because of its potential implications for the origins of humankind, evolutionary theory has been at the center of many social and religious controversies since its inception.


An early 2007 version reads

Evolution is the process in which some inherited traits in a population become more common relative to others through successive generations. This includes both pre-existing traits as well as new traits introduced by mutations. Over time, the processes of evolution can lead to speciation: the development of a new species from existing ones. All life is a result of such speciation events and thus all organisms are related by common descent from a single ancestor. [1][19]

Natural selection is a key part of this process. Since some traits or collections of traits allow an organism to survive and produce more offspring than an organism lacking them, and genes are passed on by reproduction, those that increase survival and reproductive success are more likely to be passed on in comparison to those genes that do not. Therefore, the number of organisms with these traits will tend to increase with each passing generation.[1][20][21] Given enough time, this passive process can result in varied adaptations to changing environmental conditions.[4]

Other mechanisms of evolutionary change include genetic drift, or random changes in frequency of traits (most important when the traits are, at that time, reproductively neutral), and, at the population level, immigration from other populations can bring in new traits ("gene flow") and the founder effect, in which a small group of organisms isolated from the main population will have more of the traits of the founders for many generations after isolation, even when some of the traits are detrimental.

An outline of the theory of natural selection was jointly presented to the Linnean Society of London in 1858 in separate papers by Charles Darwin and Alfred Russel Wallace. Detailed support for the theory was then set out in Darwin’s 1859 book, On the Origin of Species. In the 1930s, Darwinian natural selection was combined with the theory of Mendelian heredity to form the modern evolutionary synthesis, also known as "Neo-Darwinism". The modern synthesis describes evolution as a change in the frequency of different versions of genes, known as alleles, within a population from one generation to the next.[4] With its enormous explanatory and predictive power, this theory has become the central organizing principle of modern biology, relating directly to topics such as the origin of antibiotic resistance in bacteria, eusociality in insects, and the biodiversity of Earth's ecosystem.[22][23][24]

Which is probably nearer what we need. Adam Cuerden talk 13:25, 7 April 2007 (UTC)Reply

Thank you, Adam, for continuing to fight the good fight on behalf of accessibility. Yes, the current lead is a mess. I'd be in favor of restoring the old one (the last one you pasted) wholesale and then working from there.
A question: what did the lead look like when this article qualified as an FA?--EveRickert 00:08, 11 April 2007 (UTC)Reply

Well, bloody hell, now it's nearly gone entirely. For what it's worth, this was the lead when it was an FA. It's not perfect, but arguably better than what's there now, or was there before:

Evolution is a change in the genetic makeup of a population within a species. Since the emergence of modern genetics in the 1940s, evolution has been defined more specifically as a change in the frequency of alleles from one generation to the next. The word "evolution" is often used as a shorthand for the modern theory of evolution of species based upon Charles Darwin's theory of natural selection, which states that all modern species are the products of an extensive process that began over three billion years ago with simple single-celled organisms, and Gregor Mendel's theory of genetics. As the theory of evolution by natural selection and genetics has become universally accepted in the scientific community, it has replaced other explanations including creationism and Lamarckism. Skeptics, often creationists, sometimes deride evolution as "just a theory" in an attempt to characterize it as an arbitrary choice and degrade its claims to truth. Such criticism overlooks the scientifically-accepted use of the word "theory" to mean a falsifiable and well-supported hypothesis.

--EveRickert 20:53, 11 April 2007 (UTC)Reply

Honestly, the lead section when this article was an FA was quite lacking. It's better than what was there before, but why settle for the lesser of two evils when we can have a good? There's no reason we can't have a lead section that very briefly goes over the most essential aspects for someone who has absolutely no understanding of evolution, and then have the rest of the article go into things in more detail; although I agree that functionality is more important than blind adherence to arbitrary standards or conventions, there is a very important practical (and thus functional) reason for an article's lead section to be as short as reasonably possible: accessibility.
Evolution is a complex topic, so none of us should be surprised to see some extremely important topics covered too little, or not at all, in the lead section; that is not only tolerable, but preferable, because it means that the lead section isn't bloated. The other main concern, then, is that the lead section be reader-friendly and, in particular, informative. This involves a difficult balancing act, but there's no reason we can't reach that point of equilibrium again; we've come very close in the past.
For example, we may want to mention and link to "gene" at the start of the first paragraph, if only to account for the many uses of the word "genetic" that are simply unavoidable in the lead section to an Evolution article; however, we might not, on the other hand, need to mention DNA quite yet, and saving that for slightly later in the article will also spare us the difficulty of having to waste valuable lead space on footnotes (in the context of evolution, at least) like RNA. -Silence 21:11, 11 April 2007 (UTC)Reply

Spin-off

I think it would be really great if someone created a "Theory of evolution" article using the information already in this article. The idea would be to discuss the ins and outs of the modern synthesis and current research in greater detail without bogging down the Evolution article. For example, the lead we currently have, which is far too detailed for the average reader of this article, would work very nicely in "Theory of evolution." It would also be a great place to discuss issues like horizontal gene transfer, population bottlenecking, etc., which frankly aren't too interesting to the average reader. There's more than enough material here to make a good start on what could be a very interesting new addition to Wikipedia. Just like "Misunderstandings" spawned its own article, I think the time has come for "Theory of evolution" or "Evolutionary theory." Gnixon 13:03, 7 April 2007 (UTC)Reply

Discussions with an editor who objects to the idea.
The following discussion has been closed. Please do not modify it.
NO. There is already an easy to read article, Introduction to evolution. And second there is no "theory" of Evolution. It is a fact of Evolution. Orangemarlin 14:07, 7 April 2007 (UTC)Reply
How does a less technical article negate the usefulness of a more technical article??? Regarding theory vs. fact, please see the FAQ. Gnixon 15:40, 7 April 2007 (UTC)Reply
For comparison, see Quantum mechanics. Although there is also Introduction to quantum mechanics, the main article manages to stay nicely general, with a good balance between the theory, its applications, its history, its relationship to other fields, and its philosophical consequences. You don't see that article going on about details of, say, the WKB approximation (think HGT), and major subtopics like quantum field theory are discussed briefly and have main articles (think genetics). Gnixon 15:48, 7 April 2007 (UTC)Reply
Regarding theory vs fact, please see Evolution as theory and fact. Also see Theory of Evolution]. "Theory" of evolution is a canard to throw off the casual reader that somehow Evolution is just some random thought that entered Darwin's brain.Orangemarlin 16:52, 7 April 2007 (UTC)Reply
Give me a break. I'm talking about creating a new article that can discuss the theory in more detail, which I think would be useful. Is Theory of relativity a canard to throw off the casual reader about its truth? I'm starting to feel my temperature rising. Gnixon 17:12, 7 April 2007 (UTC)Reply
Regarding the differences with Quantum mechanics. There had been numerous conversations about simplifying this article. Those conversations (cough cough) were much easier to find a while ago, until a certain someone (cough cough) decided to reformat this discussion without consensus (cough cough). OK, I'm being passive aggressive and getting off point. Evolution is complicated, and to simplify it demeans the subject. We try to spin off forks to more easily explain certain complications. But my biggest criticism of what you write is your assumption that people are either too stupid or too lazy to read this type of article. Once again, if they want the real FACT of Evolution read this article. If they want the simplistic form, then go to the easier article. As a suggestion, why don't you suggest some changes to this article to clean up language. But do it on this page, don't mess with the main article. Maybe we can compromise between too complex and too basic. Orangemarlin 16:52, 7 April 2007 (UTC)Reply
You've implied before that I somehow messed up the archives, which I've explained before is not the case. If you're objecting to the hat/habbing, why don't we discuss it in the topic above created explicitly for that purpose, where you've so far left no comment. Gnixon 17:12, 7 April 2007 (UTC)Reply
Please stop trying to provoke a theory vs. fact debate. We usually hide those discussions on this page and refer people to the FAQ. Gnixon 17:12, 7 April 2007 (UTC)Reply
I think you're implying that the true motive of my suggestion is to allow us to dumb down this article to the level of Introduction to evolution. That's not the case. This isn't black or white. There can be a general article, a simpler one, and a more complicated, technical one. I think the latter would be very useful, and I'm not sure what your problem is with it. Gnixon 17:12, 7 April 2007 (UTC)Reply
I most certainly will "mess with the main article" if I think I can make an improvement consistent with consensus established here. WP:BB and WP:OWN are well-established policy, and good policy, to boot. I'm not saying I won't first discuss changes that might be controversial, but I really hate the idea of discouraging editors from making any changes that aren't first discussed. Gnixon 18:52, 7 April 2007 (UTC)Reply
By the way, I read the FAQ long ago, and just now. What's your point? Orangemarlin 16:54, 7 April 2007 (UTC)Reply
Evolution is both a theory and a fact. See /FAQ#Why is evolution described as though it's a fact? Isn't evolution just a theory?. I would have thought you'd be familiar with that point. Gnixon 17:12, 7 April 2007 (UTC)Reply
I'm going to ignore the callous violations of WP:CIVIL, and basically hope that you come to your senses, and try to understand what I wrote. Orangemarlin 00:55, 8 April 2007 (UTC)Reply
You know what, I actually give up. You win. You own all of these articles, because frankly, it's not worth the time and energy having an intelligent discussion with you, because you are absolutely certain of your being right on every issue. There are many more editors out there who will stand up to you, but maybe they've given up. I've never seen an editor of your intelligence level who believes in absolutes as much as you do. Go ahead and edit away, make them into Christian POV articles, if that's your wish. I'm sure you'll just archive this so you can hide what one editor feels about your editing. I'm done with you and standing up to your "I'm right and the rest of you are wrong" attitude. Enjoy buddy. 01:04, 8 April 2007 (UTC)
Paranoia Gnixon 01:39, 8 April 2007 (UTC)Reply

Sorry, Gnixon, I wasn't arround earlier to show support - of course Orangemerlin is just a troll, better to ignore. I wanted to say this: I believe that many years ago we actually had an article, theory of evolution. I think it ended up getting merged with either Natural Selection (an obvious mistake, but because at that time theory of evolution really was about Darwin's theory of evolution not the modern synthesis) or it was merged into this article. Of course, evolution is both a fact and a theory (and perhaps we should even say so in the first paragraph). In any event, content forking when an article gets long and unweildly is common, and it is not at all the same thing as simplifying an article. Slrubenstein | Talk 13:38, 9 April 2007 (UTC)Reply

Spin-off section 2

Back to the issue. What do people think of creating a more technical "Theory of Evolution"? (I'm a little disappointed that discussion on this article seems to have died down over the last couple weeks. Maybe it's the level of drama? I hope we can return to active, productive discussion---please let me know if I can somehow improve my role in it.) Gnixon 01:39, 8 April 2007 (UTC)Reply

I wouldn't support having a separate 'Theory of Evolution' article. If the lead is having problems we should fix the lead, not just split off another article. The lead of Evolution was too technical around January 1, it got better till about March 1, and recently it became too technical again. I'd also support moving more technical material to subarticles, e.g. stuff about specific genetic mechanisms. Here are some topics that, while intriguing, might not need to be covered in our main article on evolution:

  1. DNA methylation
  2. Gene flow
  3. Epigenetics
  4. Non-DNA forms of heritable variation
  5. Transposons
  6. Hill-Robertson effect
  7. Muller's ratchet

Others may have their own suggestions for what's not needed in the main article. EdJohnston 01:57, 8 April 2007 (UTC)Reply

Agree with the idea of moving things like that to main articles. (For the record, I wasn't proposing the other article only in order to move stuff from here. I really think it would be helpful to have a more technical article in one place, even if this one didn't change.) Gnixon 02:00, 8 April 2007 (UTC)Reply
I think it is a great idea to move the topical material listed above to articles on those topics, however I don't like the idea of having multiple tiers of articles on the same topics. Two is plenty, possibly too many, we should not have three. My two cents... --TeaDrinker 02:04, 8 April 2007 (UTC)Reply
I think we need Gene flow, and a brief mention of epigenetics. The others, well, the last two might be useful in explaining other things, but not more than a sentence each. Adam Cuerden talk 02:53, 8 April 2007 (UTC)Reply
I think that anything that is an important component of the theory of evolution ought to be mentioned here, with a link. But I see no reason why we canot have three tiers of articles: at the top, an article on evolution as fact and as theory that provides a general overview; then an article on the theory of evolution that goes into details about models for evolution, how they have changed, points of contention (comparable articles at this "level" would be evidence for evolution as well as articles on the evolution of actual species e.g. human evolution); then linked articles on natural selection, genetic drift, and other, more technical or contentions elements at play in current models/theorizing of evolution. Slrubenstein | Talk 13:38, 9 April 2007 (UTC)Reply

A suggestion: since it seems you really want the article to be about the modern synthesis, why not call it "Modern Synthesis" and have "Theory of Evolution" redirect there? Hey look, there is already a modern synthesis article. Maybe instead of creating a new article, you could work on this one?--EveRickert 00:04, 11 April 2007 (UTC)Reply

That's a reasonable suggestion, but Modern Synthesis, at least in its current incarnation, seems to be specifically about the historical merging of Darwin and Mendel. I was thinking of something more general that, as Slrubenstein described above, would discuss various models and aspects of the modern theory. Gnixon 21:55, 11 April 2007 (UTC)Reply

More comments from Mandaclair

Click here to expand.
The following discussion has been closed. Please do not modify it.


Leapin' Lizards, I step away for a week and things go completely bananas around here. Would it be incorrect to conclude that a Wikipedia article is *NEVER* finished, and *NEVER* satisfies everybody, and thus will *ETERNALLY* evolve and be the subject of endless tweaking and debate, as long as there is someone out there who doesn't like the way something is worded? Would it be fair to point out that in the history of writing and publishing, there has never been such thing as a board of self-appointed editors (who do not need to be experts on the topic), who can truly agree on a "consensus"? This is the "too many cooks" problem that I have already posted enough about... and it is why no other successful publishing process works like Wikipedia... But anyway:

(See WP:PERFECT. This site is what it is. Gnixon 21:02, 9 April 2007 (UTC))Reply
Sure, I get it -- but it has gotten a ton of justified criticism for that, which I really wish Wikipedia'ers would think about and try to address. If nothing else, Wikipedia sure seems like a gigantic waste of time by everybody who has 2 cents and unlimited time to contribute to a continuously evolving article that is never left to experts in the field, and is also never "done". This Evolution article has been rewritten and modified about 10,000 more times that the number of editions that the Origin of Species went through. I can't imagine that any of that is *intellectually* justified. Concepts in the field of Evolution are not changing at a rate that justifies the rate of change to this article. Like the Malagasy proverb goes: "too many people are like eels, squirming about in the mud". Mandaclair 21:32, 9 April 2007 (UTC)Reply
True, many experts leave in frustration. This article is constantly challenged and changes like a yo-yo. I partially agree with you concerns, but I should point out that many experts have strong POV's that slant an article (my understanding is that Race and Intelligence was written by an expert and it has a definite slant). The Nucleus article was written and developed by a non-expert with the assistance of experts and it turned out quite nice (well the last I looked at it, but since then who knows). I do agree that it would be nice to have some form of hard copy less prone to change. GetAgrippa 14:31, 11 April 2007 (UTC)Reply

I only want to mention that I will be working on the Speciation section a bit, and then I figure I'll leave the rest of this process to those who are more passionate about "the process". I do have two general suggestions though, about recent conversations:

1.) The desire to keep an article's lead "between four and seven paragraphs" or some other benchmark number set by Wikipedia seems absurd to me, as if both Silly Putty and the science of Evolution should be given equal lead lengths. Lousy, senseless standards pave the way for lousy, senseless writing. Do not fear a longer, more involved lead -- Evolutionary Biology is certainly worthy of it.

Standards aren't set in stone, but they usually arise for good reasons. Surely EB isn't the only complex topic Wikipedia has ever had to deal with? Gnixon 21:07, 9 April 2007 (UTC)Reply
Like evolutionary "adaptations", standards *might* arise for good reasons... or they may be baggage left over from conditions that are no longer present. I still argue -- as I teach in my courses -- that Evolutionary Biology is one of the most complex sciences *ever*, if not *the* most complex science based on what we can actually observe on Earth. Thus if any scientific topic is worthy of a bit more robust article, I think Evolution has to be it. Mandaclair 21:32, 9 April 2007 (UTC)Reply
"I teach in my courses -- that Evolutionary Biology is one of the most complex sciences *ever*, if not *the* most complex science based on what we can actually observe on Earth." Wow! POV pushing or naive of other biological disciplines. I have never published in field of evolution, but I have in neuro, cancer-immuno, developmental, and cell signalling. All these fields are changing and have drastically changed in the last twenty years and are just as complex as evolution.
I have to disagree. I publish in genomics and neuro as well as evolution, and I invite you to think of this: for every complex system in Biology you may decide to study, be it neuro, development, oncology, cell signalling, WHATEVER -- the field is made immeasureably more complex when you admit and begin studying the millions or hundreds of millions of years of evolutionary change and history that formed those systems that, traditionally, have been studied "outside" of an evolutionary point of view. But Evolution is, by all accounts, the central organizing theme of all Biology now (and isn't this in the Wikipedia article too?) -- and that is really what I mean. No level of nobel-prize winning medical or cellular inquiry can ever reach the complexity of investigating how that system came about as a result of hundreds of millions of years of history, natural selection, genetic processes, speciation, extinctions, gene and genome duplications, et cetera et cetera. Mandaclair 22:42, 10 April 2007 (UTC)Reply
It is a silly argument as it is all complex and all interlinking and related (it is still a POV unless you can provide a peer reviewed article to justify such a claim). I can't imagine any biologist not being interested in evolution (so I gather the gist of what you are saying). Further we shouldn't mention any experience or expertise (I did sorry) because without a real name to validate it can all be a bloated load (many real scientist are reluctant to post their names, and it is obvious why). Apparently numerous editors have lied about their experience and expertise so I am wary of the "good faith" clause of Wikipedia. This is not an accusation (you are obviously knowledge about the subject) but in this Wiki I encourage editors to make a case with referenced materials. Initially all my arguments were just a posit and numerous references to justify the argument, however it is amazing how many people will just ignore the literature and continue arguing POV without any backing. I swore off Wikipedia about four or five times because of the frustration of evolution related articles. Funny I completely altered one non-related science article without a single citation or any references. It is all accurate and I can provide the literature but no one has challenged or seems concerned. I firmly believe that if Pennisi, Mayr, Gould, and Dawkins could have collaborated to write this article many editors would still hate it. GetAgrippa 14:15, 11 April 2007 (UTC)Reply


If evolution can only be explained by a War in Peace length article or like Structure of Evolution theory then this is a lost cause for an encyclopedia.

A War in Peace length article is not necessary, and this is an absurd comparison. The arguments above seem to be AGAINST a lead that is "too long" based on some arbitrary measure, or an article that is "too complex" for the general nonbiological mind. Nobody wants to post a lengthy War and Peace tome here, but you simply cannot achieve transparency by sacrificing accuracy and completeness. Raw length and jargon don't need to enter into the article, but accuracy and completeness do. I'll repeat: there is no functional bubblegum summary of Evolution that is accurate and complete. Sorry. Mandaclair 22:42, 10 April 2007 (UTC)Reply
Well I apologize for the dramatic use of words. I have generally argued for disregarding concerns over the length (because I like the idea of a comprehensive article) and would prefer an article that just discussed evolution and not lots of side issues (there are so many related articles already developed to some degree that cover many topics). I have also argued to include examples of speciation in birds, fish, insects, plants, etc (pictures would be nice). I also like the idea of basic and advanced sections for articles so that the inquiring mind can pursue the topic further and in more detail. This article has no central organizing strategy for presenting the material nor any guidelines as what to present. I have argued that the correct terminology should be used but many editors see it as jargon and decreasing accessibility. I can't see an evolution definition not straightforward mentioning shifts in gene allele frequencies. One of my early comments was to develop population genetics and modeling maybe mention Hardy-Weinberg equilibrium but that was received like a Ledzepplin. Few editors suggest a central organizing plan (Silence did with strict Wiki guidelines as a rule)perhaps if you have time you could offer some suggestons. It seems a shame not to use the extensive literature to paint a rich, colorful, and expansive picture of evolution. GetAgrippa 04:17, 11 April 2007 (UTC)Reply


There are many scientist, students in field of evolution, and actual published evolutionary biologists who have contributed to this article (even the experts don't always agree on any particular point). I would agree that the conflation of novice, expert, and vandals tends to generate an ever changing article that always seems to lack focus and be somewhat disjointed.GetAgrippa 17:29, 10 April 2007 (UTC)Reply

2.) The complaints about the article not being transparent enough to "non-biology majors" are unfortunate, but I would like to argue that nothing can be done about this. Evolutionary science *IS* a complex science -- arguably the most complex science in biology -- and thus it necessarily requires a sound understanding of many concepts (yes, including genetic drift). People who argue against Evolutionary Science mainly argue against it out of sheer ignorance of the core concepts. Thus, failing to provide those concepts in their entirety will only serve to perpetuate a senseless debate. There is no way to distill evolutionary science down into a bubblegum version that everyone can understand and reconcile with their pre-existing beliefs about science and origins, (just as there is no way to distill general relativity into a pop-science version), and I would strongly argue that any attempt to write a solid, accurate, and informative article about Evolution that is accessible to "the uneducated masses" (i.e. those who are afraid of, or unfamiliar with biological concepts) -- will ultimately fail. Thanks, Mandaclair 20:52, 9 April 2007 (UTC)Reply

If you teach an introductory biology course then you are aware that some of what you teach is not current or just incorrect, but it is a starting point and a foundation for further details later. Introductory courses often teach the basics and are not current. The article should be gauged for an encyclopedia audience which is generally considered high schoolers. I have never known any academic to suppport the use of Wikipedia for their students. There is always a compromise of being concise and precise in an encyclopedia article. GetAgrippa 17:29, 10 April 2007 (UTC)Reply
Agreed... my only point is that perhaps this article has lost a lot of its accuracy (a thing different from precision) via a push to be too concise. Mandaclair 22:42, 10 April 2007 (UTC)Reply
The article need not be written to the lowest common denominator, but we should also keep in mind that this is an encyclopedia, not a textbook or journal review article, so it's important to consider the audience. If Quantum mechanics can be made accessible, surely Evolution can, too. What do you think about the idea of creating an explicitly technical article called "Theory of evolution" or something? (Please contribute to the discussion above.) Gnixon 21:07, 9 April 2007 (UTC)Reply
I would be very interested to know if working quantum physicists approve of the "accessible" Quantum Mechanics article, or if they have also abandoned it hopelessly to the editorial wolves of the popular voice. Accessible does not necessarily mean accurate, scientific, or well-written. Mandaclair 21:32, 9 April 2007 (UTC)Reply
At least one working ("quantum") physicist considers the early sections of Quantum mechanics to be sufficiently accurate and scientific and does not think the accessible language used in the article damages its accuracy. I wouldn't go too far pushing that article as a model of good Wikipedia output, but how about today's featured article, Solar System? Gnixon 21:48, 9 April 2007 (UTC)Reply
I'm not sure that a comprehension of the solar system requires the same complexity of scientific understanding that Evolution does. Even the most rabid creationist will only debate the earliest origins of the solar system, but will not argue the order of the planets, their orbital and rotational periods, their chemical compositions, etc. The comparison isn't really apt, in my opinion. Mandaclair 22:00, 9 April 2007 (UTC)Reply
Simply put: my point is that the lamentably idiotic and life-sucking popular debate against Evolutionary Science is the result of two things: 1.) Humans evolved to be more tribal than intellectual (selection for cooperation and gregariousness was stronger than selection for analytical reasoning) and therefore, conforming with your peers (political, cultural, municipal, or religious) will always prevail over intellect and reason.... and, 2.) Exactly zero percent of the individuals who argue against Evolution understand Evolution enough to argue about it intelligently. Thus if the goal of this Wikipedia article is to be educational, functional, informative, or useful *at all*, it must be complete and true to the core concepts of the Science -- accessible, or not. Mandaclair 21:40, 9 April 2007 (UTC)Reply
Hmm. I disagree with parts of both points, but I'm more concerned by the tone of disinterest in outreach from a science professor. No offense intended. Gnixon 21:52, 9 April 2007 (UTC)Reply
No offense taken, because on the contrary: I do a tremendous amount of educational outreach, more than the majority of my professional peers. However, I am generally not faced with a horde of individuals outside of my field, all jostling and competing for equal voice, authority, and editorial status. A central concept of outreach is the distinction between the educators, and those who are there to learn something. Certainly education goes both ways, but academics simply *do not* have the time for the kind of bureaucratic "education by committee" that seems to go on around here. It doesn't have anything to do with a disinterest in outreach, it is more a question of organized, accurate information vs. a chaotic editorial process. Mandaclair 22:00, 9 April 2007 (UTC)\Reply
I agree humans are not intelligent creatures but intuitive. However the same "logic and reason" of modern man has been around 100-200,000years and given rise to both religion and science. Both have evolved and are still evolving and both have had a dramatic impact on the life history of humans. The fact is both have been completely wrong at times, but they are different domains and this is not a comparison. GetAgrippa 17:29, 10 April 2007 (UTC)Reply
Absolutely! But come on man: there's NO WAY IN HELL you'll ever get this crowd of editors to allow "non-overlapping magisteria" in this article!!! "Too jargony", "Too war and peace", "Too catering to creationists", "Too opaque", etc. etc. I personally would love to see that point addressed, but I suspect there is another article somewhere that handles it. Mandaclair 22:46, 10 April 2007 (UTC)Reply
It's here. But a link to it would probably be appropriate in Creation-evolution_controversy and a few other places.--EveRickert 23:52, 10 April 2007 (UTC)Reply
Well, we seem to be conflating distinct issues. I'm not sure what the "eels in the mud" have to do with using accessible language appropriate for the readers of an encyclopedia. As for the eels, sure they can be a pain, but I don't know if they're worth getting quite so worked up about. Less patient editors should be able to find plenty of less popular and contentious articles to contribute to. For example, horizontal gene transfer and Hell on Wheels (doesn't yet exist) would probably welcome all the expert edits they can get. Gnixon 22:33, 9 April 2007 (UTC)Reply
I'm not much of an expert on horizontal gene transfer, and so I have the sense to stay away from that article. Read that sentence again.  :) Also, I am much more interested in this Hell on Wheels article, as I personally am much more involved in it, as you may already know. Mandaclair 22:41, 9 April 2007 (UTC)Reply
And see: "disagreement" with scientific points that are made is precisely a gigantic part of the editorial problem here. Whether or not you personally "disagree" that selection was stronger for cooperation and gregariousness than for rational/logical human thought, the fact remains that hominids and the great ape sister-groups were gregarious and cooperative animals for millions of years before logical thought was refined, indicating a history of very strong selection for gregariousness (which I colloquially like to call "tribalism" -- not that it necessarily needs to have anything to do with "tribes".) Throughout the history of humankind (and our primate relatives), we have done and believed countless stupid, irrational, illogical, and ridiculous things, and not gone extinct because of it... but we have always been gregarious. Evolution didn't have to go that way in humans, but it did. Compare cephalopods for an example of a lineage where intellect is likely to have been selected for more strongly than gregariousness. An interesting book on this topic is Jared Diamond's Guns, Germs, and Steel, which argues (in part) that humans were only able to indulge in philosophy, science, and the development of technology, arts, and highly rational thought, after food surpluses accumulated as a result of sedentary agricultural lifestyles. Which requires gregariousness. Mandaclair 22:23, 9 April 2007 (UTC)Reply
Give me some credit---I was disagreeing with your conclusion. We seem to be getting off-topic. Maybe we should take further discussion to our user talk pages. (I love GGS by JD. His other, nice, but not as thrilling.)Gnixon 22:33, 9 April 2007 (UTC)Reply

_____

I just saw the following comment by the now-departed Orangemarlin: "Evolution is complicated, and to simplify it demeans the subject. We try to spin off forks to more easily explain certain complications. But my biggest criticism of what you write is your assumption that people are either too stupid or too lazy to read this type of article. Once again, if they want the real FACT of Evolution read this article."

To that, I say HEAR, HERE. I am glad to see that there was another editor who took this point of view (and I don't mean POV). Too bad he was also driven away by the frustrating environment around here. It's enough to make one want to scoop one's own eyes out with a spoon.

I am currently pondering whether perhaps this group of editors may have a particular problem with anti-elitism, which is the most surefire way to drive off individuals who often have the most to contribute. A comment on my talk page, "Surely non-experts can contribute to articles in some ways and experts don't need to have their holy authority worshipped at every turn?" is the sort of comment that *Never* occurs in academic settings. My answer to that question, by the way, is generally NO. Non-experts are rarely as equipped with sufficient knowledge and experience to write the most accurate and representative articles on things. Sorry for the reality check, but that's why none of us is likely to be offered an authoring deal for a textbook or encyclopedia entry on resuable spacecraft engineering. Wikipedia is not journalism, and neither are other encyclopedias. The only thing I'll add is that "authority" is not holy and need not be worshipped, but a lot of progress might be made around here (and on Wikipedia in general) if people knew their limits, knew what they are (and are not) qualified to write about, and do not worship academic authority, but at least respect it.

I see that this particular user has driven away another experienced editor recently, with his impossible attitudes and rhetoric. I encourage the rest of you, strongly, to do something about this. Meanwhile, I'm going to make some edits to Speciation in the next couple of days, and then give up on this process in favor of more pressing (and productive) matters. It's way too much work and wasted time, for way too little progress. I'm sorry if that sounds like a poor attitude about things, but it's a very prominent one (regarding Wikipedia), and it is certainly well-justified. Mandaclair 23:00, 10 April 2007 (UTC)Reply

Wow. I think that's all I can say in a civil tone. Gnixon 00:59, 11 April 2007 (UTC)Reply
Yes, I initially thought "wow" myself when I saw you had posted that "experts don't need to have their holy authority worshipped at every turn". I guess I had wrongly assumed this was an intellectual environment, but that statement of yours just completely blows my mind, and all intellectual sensibilities out of the water. "Wow", indeed. If global educational and scientific communities operated under that philosophy, Gnixon, the human race would be nothing today but a writhing sea of murderously competitive cannibals living in war and filth -- and that is not an evolutionary hypothesis, it is a social one. Please think about what a stupid statement that was to make, in the context of what we are trying to do here as intelligent adults writing an article on science. Mandaclair 01:40, 11 April 2007 (UTC)Reply
No need to call me stupid. I suggest we close this discussion. Gnixon 01:49, 11 April 2007 (UTC)Reply
Once again: a thoroughly nonproductive, nonintellectual statement from your end, better suited for a wrestling ring, a soap opera script, or a roller derby game, than for any form of academic exchange. Nobody called *YOU* stupid. The statement you posted to my user talk, however, was incredibly and astoundingly stupid, and I will stand firmly by that. Mandaclair 02:33, 11 April 2007 (UTC) I agree to close this discussion, as I would rather spend time on the article.Reply

Does someone need to separate you two?--EveRickert 02:14, 11 April 2007 (UTC)Reply

I think I'm done with the article for now (I know, I've said that before, but at this point I've really made almost all of the edits I felt were necessary -- finally got to the Speciation and Evidence sections). In the process I've gotten a lot of backlash for what people think is an impatient, arrogant, and dismissive attitude on my part. Maybe that is justified, maybe not, but if you're curious on my true point of view on those topics (and the recent history of the actual article), please have a look at my talk page. Thanks and I'll check back in again, one of these days... probably sooner rather than later :) Oh, and P.S. I am probably changing my username to TxMCJ. Not trying to be anonymous (y'all know who I am) but I'd like to cut down on some of the user-Googling, if you catch my drift. Thanks,Mandaclair 17:50, 11 April 2007 (UTC)Reply

New lead

New lead: overly reductionist? In general I often applaud serious, bold attempts at massive streamlining, but I wonder if the lead edit by Silence isn't a bit extreme? Plus -- throwing the word theory out front so soon, in such a short lead, could cause all kinds of problems to arise due to the popular misundestanding of the word "theory".

I'm tempted to revert, but I won't "own" this article... I think many of the bold deletions Silence made might be able to really simplify the lead, but the currently posted solution might be a bit overboard... we'd also need to make sure that all of that material gets re-integrated SOMEWHERE in the article, if not in the lead. It may be detailed information, but it's not trivial information. Also: I will differ on the claim that non-organisms are non-biological. DNA is not an organism, but it is biological. As is a virus. Mandaclair 21:06, 11 April 2007 (UTC)Reply

I think that the new rewritten lead of Silence is an immense improvement. I could not believe how horribly the lead had deteriorated over the last few months. I think there is no problem with jamming technical material into the body of the article, but since the lead is probably all that over 90% of the readers will ever read, it better be well written. The lead should be short and succinct and interesting. It should not be overly technical but should give a rough idea about the subject matter. If there is material that you feel ABSOLUTELY must be included, put it in the body, not in the lead. Leave the current lead alone.--Filll 21:16, 11 April 2007 (UTC)Reply
To avoid confusion, it should be noted that Mandaclair was responding to my initial edit (which just trimmed some of the trivia out of the lead section to make it easier to see what was crucial before expanding upon that), whereas Filll is responding to my (provisional) rewrite. I don't agree with Filll that we should "leave the current lead alone"—there are a lot of improvements to be made to it, and immediately after any major change we should expect plenty of discussion and revision. However, I agree with your point that most important topics in evolution shouldn't even be alluded to in the lead section, simply because there are so many dozens of them that it would overburden our readers, plus most of them are too technical to meaningfully explain in only a few words. Objective "importance" is not the only criterion for coverage in the lead section, nor even the most important one; practical value to completely uninformed readers is. -Silence 21:30, 11 April 2007 (UTC)Reply
Thanks for a much needed and very well done edit. Agree material lost should be covered somewhere in the article. Of course this version of the lead is open to improvement, but next time we find the lead spiraling out of control, I suggest returning to this very good one. Glad to see you back here, S. Gnixon 21:36, 11 April 2007 (UTC)Reply


Impact

The rewrite is indeed excellent and really accomplished a lot in one stroke. However, historic/current resistance to evolution should be tacked on near the end; perhaps after the 1st sentence in the 3rd paragraph. Without it, there is absolutely no sense evolution was a revolutionary paradigm shift... which reminds me, that paradigm shift also needs mentioned and wikilinked (what did evolution displace). - RoyBoy 800 21:34, 11 April 2007 (UTC)Reply

Mentioning controversy in the lead may or may not be a good idea in principle, but I'm worried that it will just become a big target for warring with creationists. The last sentence has nice wording about how important evolution is to biology, and the Controversy section displays prominently in the TOC. If we mention controversy in the lead, let's be very careful about it. Gnixon 21:40, 11 April 2007 (UTC)Reply
You are talking to the guy who did the Abortion lead; not to boast or anything, but that beats Evolution hands down in the controversy category. :"D RoyBoy 800 21:56, 11 April 2007 (UTC)Reply
Haha, duly noted, but there's a lot more of substance about evolution that doesn't pertain to controversy and competes with it for space in the lead. That's to say, abortion is a relatively simple thing to describe, but its controversy is highly notable. Evolution has a somewhat smaller degree of controversy, and evolution itself is much broader and more complex. Gnixon 22:13, 11 April 2007 (UTC)Reply
True :'D, but the quality science smackdown by Silence makes me not too concerned with that anymore. The science could bloat again if it wanted to, but if controversy/social aspects are kept in a paragraph on their own, so it can be compartmentalized successfully. Just as we did for abortion, people have been killed and clinics bombed, but we kept the second paragraph down to one sentence and well placed wikilinks. - RoyBoy 800 22:20, 11 April 2007 (UTC)Reply
I don't see the need for such a tack-on, RoyBoy, as this is a biological article, rather than a sociological or historical one. The top priority of this article is to explain to our readers the scientific understanding of biological evolution; whether or not it constituted a "paradigm shift" (itself a somewhat controversial idea within philosophy of science; it would probably be opening an unnecessary can of worms for Wikipedia to endorse a specific perspective on it here!) is at best an afterthought, and arguably barely merits inclusion in the article body (perhaps in the "Social effect" and/or "History" section), much less in the lead. Remember that at the end of the lead section we haven't even begun explaining many of the basics of what is actually physically happening in evolution; compared to that, evaluations of its social significance are, at least for the purposes of a biology article like this, of peripheral importance.
I also doubt that one sentence could properly convey the idea without misrepresenting the scope and significance of the controversy—especially since this would be the only sentence in the lead section not dealing directly or indirectly with the science of evolution. Describing "resistance" in such a context would imply that there is significant scientific resistance to evolutionary theory, which couldn't be further from the truth. Furthermore, I would like to keep the third paragraph as short as possible, and expand the "History" section instead where possible, because that section is currently woefully diminished. -Silence 21:43, 11 April 2007 (UTC)Reply
Top priority does not denote only priority, for an article, and especially for the lead. I hate to be the spoiled sport, but I must remind everyone here this is not a biology article. Okay? This is an article on the subject of evolution. This includes biology and controversy. No mention of controversy is a glaring oversight. See the Encarta beginning for a guildeline. Evolution displaced dominant historic views; if that isn't lead material, I don't know what is. The focus should, is and always has been on the biology. Great! That does not give us license to push other stuff to the bottom of the article. - RoyBoy 800 21:54, 11 April 2007 (UTC)Reply
This is an article about biological evolution. (There are other articles for other types of evolution.) Thus, it is a biology article. Controversy is relevant here only insofar as it is relevant to understanding biological evolution. No mention of controversy in the lead section is infinitely less of an oversight than no mention of dozens of other, more important topics for understanding biological evolution, like the fossil record and DNA. Yet these, too, are mentioned nowhere in the lead. For an article as immensely complex and broad as this one, we simply need to stop trying to squeeze every single "important" topic into the lead section, or it'll grow unmanageably large once again; if something is "important" we should work first on improving its coverage in the article body, and only afterwards, if there is wide agreement, insert it into the lead. If anything even the current lead is a little longer than would be ideal.
Every major scientific discovery in history has "displaced dominant historic views", in one way or another. Without proper context and details, this is too vacuously vague to be very useful to readers in the "bite-sized" format that a lead section demands.
The introduction to the Encarta evolution article is twice as long as the lead section of evolution. Since that means that at least half of the information in that introduction wouldn't fit here without us beginning to re-bloat the newly-trimmed section, pointing to the inclusion of something there wouldn't be sufficient grounds for inclusion even if Encarta was the pinnacle of encyclopedic achievement.
The fact that this article's focus is on biology not only gives us "license" to push other stuff to the bottom of the article (and to other areas of the article body, many of which desperately need just that kind of "pushing" in order to flesh out missing information!); it gives us the duty to do so. -Silence 22:18, 11 April 2007 (UTC)Reply
False comparison, DNA is implicitly referred to in the lead multiple times, and fossils are simply mineralized relics of DNA variation. Heh, yeah that's a stretched connection... but the point is both are central evidence for evolution. Controversy is a different sub-topic entirely; having nothing to do with biology, but everything to do with evolution... especially historically, the history (of the controversy and evolution's historic context) are under serviced in the lead.
I get the distinct impression that now that the lead is "in shape", nothing can be added? Another way to see it, is that you've created room for other notable aspects of evolution to be mentioned.
Don't obfuscate the issue with re-bloat. Encarta is a clear example that an encyclopedic article and lead is not exclusive to its main subject. Ever. Encarta is long and I have no intention of replicating the topics/coverage it has.
The disambig notice at the top clarifies what concept of "Evolution" this article is covering. It's function is to keep people from placing concepts from Stellar evolution here. It provides absolutely no editorial mandate to focus exclusively on the biological aspect of that concept. I'll understand if I need to repeat this several times since this has obviously been an assumption carried forward by the dominant/active editors here, but that disambig notice does not change the fact this article needs cover all aspects of biological Evolution. That includes controversy (religion), history, politics and if notable enough, sociology. I'm not debating this with you, I'm trying to, with as light a touch as possible to a valuable contributor, to say... incorrect. Disambiguation is just that, disambiguation; it does not set (or force) tone on an article. - RoyBoy 800 22:52, 11 April 2007 (UTC)Reply
But aren't you underestimating the notability of the impact of biological science on society? For analogy, is it reasonable for the Physics lead to mention the impact of say, nuclear weapons and semiconductors? Gnixon 22:22, 11 April 2007 (UTC)Reply
This article isn't analogous to physics because it discusses a biological process, rather than a field or discipline of science; the proper analogy would be between physics and evolutionary biology. The social impact of a certain area of science is of more relevance to the article on the study of a phenomenon (e.g., evolutionary biology) or the explanation of that phenomenon (e.g., modern evolutionary synthesis) than on the phenomenon itself (e.g., evolution). This is not to say that the social impact of such study isn't important enough to mention in this article, merely that it's not quite important enough for the lead section, if only because there's always so much vastly more important information that we're currently leaving out for the sake of brevity and comprehensibility. -Silence 22:28, 11 April 2007 (UTC)Reply
That's what I get for picking a poor analogy. It's the idea of evolution that has impact, not the products of the field (physics). Maybe a better analogy would be Marxism or Adam Smith's take on economics. Meh. I agree with your comment below that these things should be addressed in the body before updating the intro. Gnixon 22:44, 11 April 2007 (UTC)Reply
An analogy I would refer to is Age of the Earth, which I helped out on. While almost entirely science oriented there still needs to be mention of notable dissenting views. Now here is the kicker, those views have sub-articles, young earth creationism and such... but they do still merit a mention in the parent article. As they are indeed a part, a small part, but nonetheless a part of the subject matter for the article. - RoyBoy 800 23:16, 11 April 2007 (UTC)Reply
Information that should be left out. - RoyBoy 800 22:52, 11 April 2007 (UTC)Reply
We don't have to say it was a paradigm shift, but we must provide historic context for evolution; and wikilink to the dominant scientific theory prior to Darwin. If memory serves, it Gradualism or something like that... Huxley comes to mind; it was based on slow changes of terrain being analogous to biological changes. I can't really remember. - RoyBoy 800 22:07, 11 April 2007 (UTC)Reply
I think you make a good point that this topic deserves to be broader than just the science, but it will be challenging to discuss the social impact without bloating the lead or giving undue weight to objections to evolution. Do you have specific suggestions? Gnixon 22:01, 11 April 2007 (UTC)Reply
Not this second, but it should be short, a sentence or two; although it think it could grow to a small paragraph with historic, Darwin's time, and modern sentences providing a clear understanding resistance has been notable, historic and is ongoing in certain places. - RoyBoy 800 22:07, 11 April 2007 (UTC)Reply
Such an addition might be a superb one for the exceedingly short section "History of modern evolutionary thought"; I recommend adding it there first. If something isn't even important enough for the article body, it's certainly not important enough for the lead section. Also, gradualism (proposed in 1795 by Hutton) is part of the essential basis of evolution (and of modern geology and evolutionary biology), not the "dominant scientific theory prior to Darwin"; perhaps you're thinking of Lamarckism. Regardless, none of this is remotely significant enough for the lead section. -Silence 22:23, 11 April 2007 (UTC)Reply
I agree it at least makes sense to start by including these things in the body before the lead. Gnixon 22:44, 11 April 2007 (UTC)Reply
It's not notable for a biology article, but this isn't just biology article. Again look at Abortion, we have sub-articles wikilinked in the lead; which is still very tightly written. - RoyBoy 800 22:59, 11 April 2007 (UTC)Reply

Common descent

Are we ever going to have a war about this? The lead now states common descent quite baldly as tracing down to a single ancestor. Are we resolved to ignore confusion on this subject, or should we somehow amend the sentence to clarify? Will I ever shut up about this? Graft 22:33, 11 April 2007 (UTC)Reply

If your problem is with the fact that we say "common ancestor" rather than "common ancestor or ancestral gene pool" (which will just confuse most readers, since we haven't yet said what a gene pool is), dictionary.com attests to the fact that a "common ancestor", in this context, need not be a singular, specific individual organism; it defines a "common ancestor" as "the most recent ancestral form or species from which two different species evolved". The universal common ancestor can thus be a grouping of organisms, at least as far as I can tell. That's the very reason that terms like last universal ancestor are so often used. -Silence 22:45, 11 April 2007 (UTC)Reply
Quoi? Non, as far as I've read. Maybe a population or species, as is certainly the case with "common ancestors" for sexually reproducing creatures, but certainly not a grouping of disparate organisms that are genetically distinct. Common descent should mean a single root to the tree of life. Graft 23:52, 11 April 2007 (UTC)Reply
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  3. ^ Haldane, J.B.S. (1953). "The measurement of natural selection". Proceedings of the 9th International Congress of Genetics. 1: 480–487.
  4. ^ a b c d e "Mechanisms: the processes of evolution". Understanding Evolution. University of California, Berkeley. Retrieved 2006-07-14.
  5. ^ Gould, Stephen J. (2002). The Structure of Evolutionary Theory. Belknap Press. ISBN 0-674-00613-5.
  6. ^ Dawkins, Richard (1989). The Selfish Gene. Oxford University Press Press. ISBN 0-674-00613-5.
    Lake, James A. (2004). "The Ring of Life Provides Evidence for a Genome Fusion Origin of Eukaryotes" (PDF). Nature. 431. Retrieved 2007-03-18. {{cite journal}}: Unknown parameter |coauthors= ignored (|author= suggested) (help)
  7. ^ UCLA Report (2004). "Ring of Life". Retrieved 2007-03-16. {{cite journal}}: Cite journal requires |journal= (help); Unknown parameter |month= ignored (help)
  8. ^ Doolittle, Ford W. (February 2000). "Uprooting the Tree of Life". Scientific American: pp. 72-77. {{cite journal}}: |pages= has extra text (help)CS1 maint: year (link)
  9. ^ Lake, James A. and Maria C. Riveral (1999). "Horizontal gene transfer among genomes: The complexity hypothesis". PNAS (Proceedings of the National Academy of Science). 96:7: pp. 3801-3806. Retrieved 2007-03-18. {{cite journal}}: |pages= has extra text (help)
  10. ^ Bapteste; et al. (2005). "Do Orthologous Gene Phylogenies Really Support Tree-thinking?". BMC Evolutionary Biology. 5:33. Retrieved 2007-03-18. {{cite journal}}: Explicit use of et al. in: |author= (help)
  11. ^ Gogarten, Peter (2000). "Horizontal Gene Transfer: A New Paradigm for Biology". Esalen Center for Theory and Research Conference. Retrieved 2007-03-18.
  12. ^ "IAP STATEMENT ON THE TEACHING OF EVOLUTION" (PDF). the Interacademy Panel on International Issues. Retrieved 2007-03-20.
  13. ^ "Statement on the Teaching of Evolution" (PDF). American Association for the Advancement of Science. 2006. Retrieved 2007-03-20.
  14. ^ Futuyma, Douglas J. (2005). Evolution. Sunderland, Massachusetts: Sinauer Associates, Inc. ISBN 0-87893-187-2.
  15. ^ Gould, Stephen J. (2002). The Structure of Evolutionary Theory. Belknap Press. ISBN 0-674-00613-5.
  16. ^ Lande, R. (1983). "The measurement of selection on correlated characters". Evolution. 37: 1210–1226. {{cite journal}}: Unknown parameter |coauthors= ignored (|author= suggested) (help)
  17. ^ Futuyma, Douglas J. (2005). Evolution. Sunderland, Massachusetts: Sinauer Associates, Inc. ISBN 0-87893-187-2.
  18. ^ Haldane, J.B.S. (1953). "The measurement of natural selection". Proceedings of the 9th International Congress of Genetics. 1: 480–487.
  19. ^ Gould, Stephen J. (2002). The Structure of Evolutionary Theory. Belknap Press. ISBN 0-674-00613-5.
  20. ^ Lande, R. (1983). "The measurement of selection on correlated characters". Evolution. 37: 1210–1226. {{cite journal}}: Unknown parameter |coauthors= ignored (|author= suggested) (help)
  21. ^ Haldane, J.B.S. (1953). "The measurement of natural selection". Proceedings of the 9th International Congress of Genetics. 1: 480–487.
  22. ^ Myers, PZ (2006-06-18). "Ann Coulter: No evidence for evolution?". Pharyngula. scienceblogs.com. Retrieved 2006-11-18.
  23. ^ IAP Statement on the Teaching of Evolution Joint statement issued by the national science academies of 67 countries, including the United Kingdom's Royal Society (PDF file)
  24. ^ From the American Association for the Advancement of Science, the world's largest general scientific society: 2006 Statement on the Teaching of Evolution (PDF file), AAAS Denounces Anti-Evolution Laws